We publish high quality peer-reviewed articles on animal behavior, comparative physiology, evolution, ecology, and systematics and biogeography on animals taxa from all parts of the world.
OPEN ACCESS (Free Page Charge)
Zoological Studies is an OPEN ACCESS international journal and FREE OF PAGE CHARGES from authors.

Edited by:
Benny K.K. Chan (Editor-in-Chief)
Sen-Lin Tang (Co-Editor-in-Chief)

Impact Factor: 1.257
ISI Journal Citation Reports
Ranking: 2019: 78/168 (Zoology)

ISSN (E): 1810-522X
Follow us on Twitter @ZooStudies and Facebook

Zoological Studies is listed in PubMed
Sumbit manuscript

Zoological Studies Editorial Team

The Editor in Chief, Dr. Benny K.K. Chan, and the Co-editors in Chief, Dr. Sen-Lin Tang are professors/ research fellows at Biodiversity Research Center, Academia Sinica, Taiwan.

The editors and co-editors in chief are supported by 30 Associate Editors including Drs.Sheng-Feng Shen, I-Min Tso, Pung-Pung Hwang, Yeong-Choy Kam, David Miller, Shou-Hsien Li, Chung-Pin Lin, John Wang, Claudia Castellani, Yasuyuki Hashiguchi, Chih-Ming Hung, Chiun-Cheng Ko, Chi-Chien Kuo, Teng-Chiu Lin, Jian-Nan Liu, Ryuji Machida, Hiroomi Miyamoto, Yoko Nozawa, Gray A. Williams, Benny K.K. Chan, Allen Chen, Ka Hou Chu, Jens Thorvald Høeg, Jen-Pan Huang, Hin-Kiu Mok, Felipe Polivanov Ottoni, James D. Reimer, Shen-Horn Yen, Sen-Lin Tang, and Daryi Wang.

(See Editorial Board for details)

Message from the Editor in Chief

We take up a responsibility for the academic society

    Nowadays, we often need to pay expensive page charges/open-access fees to publish our works as most of the journals are now run by business publishers. Is it possible there are quality assured, peer-reviewed and open-access SCI scientific journals, in which colleagues can publish their research achievement free? We, Biodiversity Research Center, Academia Sinica, Taiwan, attempts to take up this responsibility. We run Zoological Studies, to provide a free SCI open-access publication platform for colleagues to share out their research achievements.
    Publish articles free of page charges does not mean Zoological Studies is lack of good quality. All the accepted articles passed a formal review system. Impact factors is an indication for the article citations in a journals and has implications for journal quality. However, article citations can be also affected by the ways of marketing methods for journals in commercial publishers. We are trying hard to maintain a good impact factor for Zoological Studies. All articles are 1) formally applied for a DOI number and being open-access in the web, 2) listed and housed in PubMed database and 3) promoted in our twitters and facebook. We are not under any umbrella of commercial publishers. All the editors and reviewers are volunteers for the academics. We hope you can support us to submit your quality works to Zoological Studies, and upload your published articles in Research Gate to enhance citations as well.

Prof. Benny K.K. Chan

Recent published articles in 2020
Patterns of Bird Diversity and Endemism Along an Elevational Gradient in the Southern Mexican Highlands
Edson A. Alvarez-Alvarez, Rosalba Rodríguez-Godínez, Pablo Sierra-Morales, Sandy A. MedinaValdivia, Estefanía Vázquez-Salgado, Marlene Brito-Millán, and R. Carlos Almazán-Núñez (doi:
AbstractPDF (840KB)
Population Genetic Structure of A Marine Pelagic Egg Producer and Popular Marine Aquarium Species, the Mandarinfish Synchiropus splendidus
Priscilla T.Y. Leung, Ka Yan Ma, Min Liu, Serge Planes, and Yvonne Sadovy de Mitcheson (doi:
AbstractPDF (940KB)
Phylogeography of Triplophysa stenura (Nemacheilidae): Responded to the Mid-Pleistocene Climate Transition in the Qinghai–Tibetan Plateau
Jiaxin Hu , Mingdian Liu, and Dekui He (doi:
AbstractPDF (2.33MB)
Molecular phylogeny suggesting the single origin of Cinnamomum-associated Bruggmanniella (Diptera: Cecidomyiidae) in Asia, and Descriptions of Three New and One Newly Recorded Species from Taiwan
Sheng-Feng Lin, Man-Miao Yang, and Makoto Tokuda (doi:
AbstractPDF (1.49MB)
Seasonality and Longevity of the Functional Chloroplasts Retained by the Sacoglossan Sea Slug Plakobranchus ocellatus van Hasselt, 1824 Inhabiting a Subtropical Back Reef Off Okinawa-jima Island, Japan
Shu Chihara, Takashi Nakamura, and Euichi Hirose (doi:
AbstractPDF (0.97MB)
Taxonomic Revision and Evolutionary Phylogeography of Dusky Leaf Monkey (Trachypithecus obscurus) in Peninsular Malaysia
Nor Rahman Aifat, Muhammad Abu Bakar Abdul-Latiff, Christian Roos, and Badrul Munir Md-Zain (doi:
AbstractPDF (778KB)
Vertebrate Scavengers Control Abundance of Diarrheal-causing Bacteria in Tropical Plantations
Norman T-L Lim, Douglas A Kelt, Kelvin KP Lim, and Henry Bernard (doi:
AbstractPDF (808KB)
A New Species of Predatory Nudibranch (Gastropoda: Trinchesiidae) of the Scleractinian Coral Goniopora
Juntong Hu, Yanjie Zhang, Sam King Fung Yiu, James Yang Xie, and Jian-Wen Qiu (doi:
AbstractPDF (2.85MB)
Protandrous Hermaphroditic Reproductive System in the Adult Phases of Mothocya renardi (Bleeker, 1857) (Cymothoidae: Isopoda: Crustacea) – Light and Electron Microscopy Study
Aneesh Panakkool Thamban adn Sudha Kappalli (doi:
AbstractPDF (4.29MB)
Ecological Morphology of Neotropical Bat Wing Structures
Dennis Castillo-Figueroa (doi:
AbstractPDF (2.89MB)
Diversity of Taiwanese Brackish Crabs Genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) based on DNA Barcodes, with Descriptions of Two New Species
Jhih-Wei Hsu and Hsi-Te Shih (doi:10.6620/ZS.2020.59-59)

AbstractPDF (2.89MB)
Diversity of Bivalve Mollusks Associated with Macroalgae on the Continental Shelf of the States of Alagoas, Sergipe and Bahia, Northeastern Brazil
Lucas Santos, J. Weverton S. Souza, Silvio Lima, and Carmen Guimarães (doi:
AbstractPDF (5.09MB)
Spatial Distribution of Medusa Cunina octonaria and Frequency of Parasitic Association with Liriope tetraphylla (Cnidaria: Hydrozoa: Trachylina) in Temperate Southweatern Atlantic Waters
Francisco Alejandro Puente-Tapia, Florencia Castiglioni, Gabriela Failla Siquier, and Gabriel Genzano (doi:
AbstractPDF (8.45MB)
Ontogenetic Development Related to Parental Care of a Neotropical Fish, Pterygoplichthys ambrosettii (Siluriformes: Loricariidae)
Renato Braz de Araujo and Francisco Langeani (doi:
AbstractPDF (3.68MB)
Molecular Approach to the Identification of Three Closely Related Slugs Deroceras Species (Gastropoda: Eupulmonata: Agriolimacidae)
Kamila S. Zając and Daniel Stec (doi:
AbstractPDF (8.97MB)
Estrogen Receptor 1 (ESR1) Agonist Induces Ovarian Differentiation and Aberrant Müllerian Duct Development in the Chinese Soft-shelled Turtle, Pelodiscus sinensis
Kenji Toyota, Shoichiro Masuda, Sarina Sugita, Kaori Miyaoku, Genki Yamagishi, Hiroshi Akashi, and Shinichi Miyagawa (doi:
AbstractPDF (580KB)
Relationships and Description of a New Species of Trichomycterus (Siluriformes: Trichomycteridae) from the Rio Paraíba do Sul Basin, South-eastern Brazil
Paulo J. Vilardoa, Axel M. Katza, and Wilson J.E.M. Costaa (doi:
AbstractPDF (4.95MB)
Light Microscopic and Ultrastructural Characteristics of Heterophil Toxicity and Left Shifting in Green Sea Turtles (Chelonia mydas) from Taiwan
Yi-Fan Wang, Tsung-Hsien Li, Yi-Fan Jiang, Chau-Hwa Chi, I-Jiunn Cheng, Chiung-Hsiang Cheng, Ray Sun, and Pin-Huan Yu (doi:
AbstractPDF (5.22MB)
Pattern of Gonadal Sex Differentiation in the Rice Field Frog Hoplobatrachus rugulosus (Anura: Dicroglossidae)
Thrissawan Traijitt, Noppadon Kitana, and Jirarach Kitana (doi:
AbstractPDF (19.3MB)
Geographical Patterns in the Architecture of Neotropical Flower-visitor Networks of Hummingbirds and Insects
Leuzeny Teixeira Moreira, Luiz Alberto Dolabela Falcão, and Walter Santos de Araújo (doi:
AbstractPDF (988KB)
Accessing Multiple Paternity in the Shortfin Mako Shark (Isurus oxyrinchus)
Shang Yin Vanson Liu, Wen-Pei Tsai, Meng shan Lee, and Hsiu-Wen Chien (doi:
AbstractPDF (736KB)
Somniosus (Rhinoscymnus) cheni sp. nov., A New Species of Sleeper Shark (Squaliformes: Somniosidae) from Eastern Taiwan, with Aspects of Embryo Biology
Hua-Hsun Hsu, Chia-Yen Lin, and Shoou-Jeng Joung (doi:10.6620/ZS.2020.59-48)
Abstract PDF (16.5MB)
A New Species of the Burrowing Crayfish Genus Parastacus Huxley, 1879 (Decapoda, Parastacidae) from the Sinos River Basin, Southern Brazil
Augusto F. Huber, Emily R. Rockhill, Paula B. Araujo, and Felipe B. Ribeiro (doi:10.6620/ZS.2020.59-47)
AbstractPDF (12.3MB)
Special Issue: Fossil and Modern Clam Shrimp (Branchiopoda: Spinicaudata, Laevicaudata)
(articles 32-46) communicated by Thomas A. Hegna and D. Christopher Rogers
The World’s First Clam Shrimp Symposium: Drawing Paleontology and Biology Together
Thomas A. Hegna and D. Christopher Rogers (doi:
AbstractPDF (319KB)
Spinicaudata Catalogus (Crustacea: Branchiopoda)
D. Christopher Rogers (doi:
AbstractPDF (2.35MB)
Phylogeny and Biogeography of Spinicaudata (Crustacea: Branchiopoda)
Martin Schwentner, Nicolas Rabet, Stefan Richter, Gonzalo Giribet, Sameer Padhye, Jean-François Cart, Céline Bonillo, and D. Christopher Rogers (doi:
AbstractPDF (22MB)
The Fossil Record of the Clam Shrimp (Crustacea; Branchiopoda)
Thomas A. Hegna and Timothy I. Astrop (doi:
AbstractPDF (519KB)
A New Eulimnadia (Branchiopoda: Spinicaudata: Limnadiidae) from the US Virgin Islands
D. Christopher Rogers and Edwin Cruz-Rivera (doi:
AbstractPDF (4.12MB)
Review of the Eulimnadia (Branchiopoda: Spinicaudata: Limnadiidae) of Tropical Asia, with the Description of a New Species
La-orsri Sanoamuang, Sameer M. Padhye, and D. Christopher Rogers (doi:
AbstractPDF (4.45MB)
Description of the First Leptestheria Species (Branchipoda, Spinicaudata) from Brazil
Mélanie Van Weddingen and Nicolas Rabet (doi:
AbstractPDF (6.91MB)
Distribution of Clam Shrimps (Crustacea: Laevicaudata and Spinicaudata) in South Africa, with New Records from the Northern Cape Province
Elizabeth Meyer-Milne, Musa C. Mlambo, and D. Christopher Rogers (doi:
AbstractPDF (4.88MB)
Diagnosing Eulimnadia and Paralimnadia (Branchiopoda: Spinicaudata: Limnadiidae)
Brian V. Timms and D. Christopher Rogers (doi:
AbstractPDF (4.38MB)
Revision of Two Spinicaudatan Species from the Cañadón Asfalto Formation (Jurassic), Patagonia Argentina
Mateo D. Monferran, Oscar F. Gallego, and Nora G. Cabaleri (doi:
AbstractPDF (8.65MB)
New Spinicaudatan Species of Late Jurassic Linglongta Phase of Yanliao Biota from Western Liaoning, China
Gang Li (doi:
AbstractPDF (21.5MB)
Developmental and Functional Morphology of Eulimnadia braueriana Ishikawa, 1895 (Branchiopoda: Spinicaudata) Feeding Structures: Combination of Filtering and Scraping Feeding Mechanisms
Jer-Yuh Liu, Chun-Chieh Wang, and D. Christopher Rogers (doi:
AbstractPDF (26MB)
Testing Weissman’s Lineage Selection Model for the Maintenance of Sex: The Evolutionary Dynamics of Clam Shrimp Reproduction over Geologic Time
Timothy I. Astrop, Lisa Park Boush, and Stephen C. Weeks (doi:
AbstractPDF (4.83MB)
Population Density Effects on Carapace Growth in Clam Shrimp: Implications for Palaeontological Studies
Manja Hethke and Stephen Weeks (doi:
AbstractPDF (28.4MB)
Fossil and Modern Clam Shrimp (Branchiopoda: Spinicaudata, Laevicaudata): The World’s First Clam Shrimp Symposium and a Celebration of Brian V. Timms
D. Christopher Rogers and Thomas A. Hegna (doi:
AbstractPDF (3.88MB)
Special issue finished
Checklist and Distribution of Freshwater Cladocera (Crustacea: Branchiopoda) in Côte d’Ivoire (West Africa)
Raphaël N’doua Etilé, Théophile Aké Bédia, Georges Kassi Blahoua, Gouli Gouré Bi, Paul Essetchi Kouamelan, and Valentin N’Douba (doi:10.6620/ZS.2020.59-31)
AbstractPDF (10.5MB)
A New Species of Predatory Nudibranch (Gastropoda: Trinchesiidae) of the Coral Pavona decussata
Juntong Hu, Yanjie Zhang, James Yang Xie, and Jian-Wen Qiu (doi:10.6620/ZS.2020.59-30)
AbstractPDF (10.3MB)
Kristianides, A New Genus of Scale Worms from the Northern Gulf of Mexico and New Records of Phyllohartmania taylori (Annelida: Polychaeta: Polynoidae)
Patricia Salazar-Silva (doi:
AbstractPDF (7.66MB)
Lacydoniidae (Annelida) Off the Coast of North-eastern Japan: A Description of Lacydonia japonica sp. nov.
Natsumi Hookabe, Naoto Jimi, Shinji Tsuchida, Yoshihiro Fujiwara, and Hiroshi Kajihara (doi:
AbstractPDF (3.57MB)
What is Sesarmops impressus (H. Milne Edwards, 1837) (Crustacea: Brachyura: Sesarmidae)?
Peter K. L. Ng, Jheng-Jhang Li, and Hsi-Te Shih (doi:
AbstractPDF (88.3MB) | Erratum notice
A New Fiddler Crab of Austruca Bott, 1973, Closely Related to A. perplexa (H. Milne Edwards, 1852) (Crustacea: Brachyura: Ocypodidae), from the South Pacific Islands
Hsi-Te Shih and Joseph Poupin (doi:
AbstractPDF (27.4MB)
Simultaneous Occurrence of Diapause and Cold Hardiness in Overwintering Eggs of the Apple Oystershell Scale, Lepidosaphes Malicola Borchsenius (Hem.: Diaspididae)
Parvaneh Nazari, Nafiseh Poorjavad, and Hamzeh Izadi (doi:
AbstractPDF (2.78MB)
A New Genus and Two New Species of Freshwater Crab (Crustacea: Brachyura: Potamidae) with Unusual Coiled Tip of Male Second Gonopods from Yunnan, Southwestern China
Chao Huang, Jian Wang, and Hsi-Te Shih (doi:
AbstractPDF (9.2MB) | Erratum notice
Integrative Descriptions of Two New Macrobiotidae Species (Tardigrada: Eutardigrada:  Macrobiotoidea) from French Guiana and Malaysian Borneo
Daniel Stec, Magdalena Dudziak, and Łukasz Michalczyk (doi:10.6620/ZS.2020.59-23)
AbstractPDF (17.3MB)
Population Genetic and Social Structure Survey of Solenopsis geminata in Thailand
Mingkwan Nipitwattanaphon, Akarapong Swatdipong, Sasitorn Hasin, and John Wang (doi:10.6620/ZS.2020.59-22)
AbstractPDF (5.00MB)