Four New Species and One New Record of Land Hermit Crabs (Coenobita) from Indonesia
Four new species of land hermit crabs from Indonesia are described based on morphological differences and mitochondrial 16S rDNA and COI data. Coenobita moluccensis, C. patsyae, and C. celebensis form a major clade in the phylogenetic tree, exhibiting morphological similarities, particularly in the structure of the male sexual tubes. Coenobita granularis, from Central Sulawesi, shares morphological similarities with the genetically closely related C. pseudorugosus, especially in the male sexual tubes. In addition, a newly recorded species, C. variabilis, has been confirmed in West Papua. This study increases the total number of Coenobita species known from Indonesia to 13.

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