Article
Vol. 48-2-11, 2009
Pterostichus macrogenys Bates (Coleoptera: Carabidae) and Its Allied Species of Northern Japan: Descriptions of Seven Additional Species and Possible Evidence Supporting Species Status
Kôji Sasakawa (2009) Pterostichus macrogenys Bates (Coleoptera: Carabidae) and its allied species of northern Japan are revised based mainly on the structures of the male endophallus (inner sac everted from the aedeagus).  Seven species are newly described: P. shirakamisan sp. nov., P. takadateyamanus sp. nov., P. chokaisanus sp. nov., P. gassanus sp. nov., P. eboshiyamanus sp. nov., P. yahikosanus sp. nov., and P. iwakiensis sp. nov.  Collection data and body-size differences among these species suggest the sympatric occurrence of large and small species in some localities.  The implications of these results for the species-level taxonomy of this group are discussed.
Keywords
Ground beetle, Male endophallus, New species, Nialoe, Species status.