We publish high quality peer-reviewed articles on animal behavior, comparative physiology, evolution, ecology, and systematics and biogeography on animals taxa from all parts of the world.
OPEN ACCESS (Free Page Charge)
Zoological Studies is an OPEN ACCESS international journal and FREE OF PAGE CHARGES from authors.

Edited by:
Benny K.K. Chan (Editor-in-Chief)
Sen-Lin Tang (Co-Editor-in-Chief)

Impact Factor: 1.904
ISI Journal Citation Reports
Ranking: 2021: 62/177 (Zoology)

ISSN (E): 1810-522X
Follow us on Twitter @ZooStudies and Facebook

Zoological Studies is listed in PubMed
Sumbit manuscript

Zoological Studies Editorial Team

The Editor in Chief, Dr. Benny K.K. Chan, and the Co-editors in Chief, Dr. Sen-Lin Tang are professors/ research fellows at Biodiversity Research Center, Academia Sinica, Taiwan.

The editors and co-editors in chief are supported by 30 Associate Editors including Drs.Sheng-Feng Shen, I-Min Tso, Pung-Pung Hwang, Yeong-Choy Kam, David Miller, Shou-Hsien Li, Chung-Pin Lin, John Wang, Claudia Castellani, Yasuyuki Hashiguchi, Chih-Ming Hung, Chiun-Cheng Ko, Chi-Chien Kuo, Teng-Chiu Lin, Jian-Nan Liu, Ryuji Machida, Hiroomi Miyamoto, Yoko Nozawa, Gray A. Williams, Benny K.K. Chan, Allen Chen, Ka Hou Chu, Jens Thorvald Høeg, Jen-Pan Huang, Hin-Kiu Mok, Felipe Polivanov Ottoni, James D. Reimer, Shen-Horn Yen, Sen-Lin Tang, and Daryi Wang.

(See Editorial Board for details)

Message from the Editor in Chief

We take up a responsibility for the academic society

Nowadays, journals often require expensive page charges/open-access fees to publish scientific research articles. Is it possible there are quality assured, peer-reviewed and open-access SCI scientific journals, in which colleagues can publish their research without these hefty fees? We, the Biodiversity Research Center, Academia Sinica, Taiwan, have taken up this responsibility. We publish a journal entitled Zoological Studies, to provide a free SCI open-access publication platform for colleagues to share their research achievements.
    Publishing articles free of page charges does not mean that articles in Zoological Studies lack quality. All accepted articles have passed a formal review system. Impact factors are often used to judge the quality of a journal. However, article citations can be also affected by the  marketing methods used in journals published by commercial publishers. We expect to maintain a respectable impact factor for Zoological Studies. All articles are 1) formally assigned a DOI number and are open-access on the web, 2) listed and housed in PubMed database and 3) promoted in our twitter and facebook accounts. We are not under the umbrella of any commercial publisher. All the editors and reviewers are volunteers. We hope you can support us by submitting your quality research to Zoological Studies, and subsequently upload your published articles in Research Gate to enhance citations as well.

Prof. Benny K.K. Chan

Recent published articles in 2022
Description of A New Archaeococcoid of the Genus Perissopneumon Newstead (Hemiptera: Coccomorpha: Monophlebidae) from India
Anirban Das and Bijan Kumar Das (doi:-)
Abstract丨PDF(-MB)(Proof production in process)
Abundance Trends and Nesting Biology of Green Turtles Chelonia mydas (Testudines: Cheloniidae) During Ten Consecutive Breeding Seasons at Akyatan Beach, Turkey
Can Yılmaz, Ayşe Oruç, and Oguz Turkozan (doi:-)
Abstract丨PDF(-MB)(Proof production in process)
An Updated Checklist of Sea Slugs (Gastropoda, Heterobranchia) from Hong Kong Supported by Citizen Science
Lai Him Chow, Vriko Pik Fan Yu, Zi Yi Kho, Gomen Chun Lung See, Adam Wang, David M. Baker, and Ling Ming Tsang (doi:-)
Abstract丨PDF(-MB)(Proof production in process)
Redescription of the Pplanthopper Bursinia genei (Dufour, 1849), with a New Record from Greece (Hemiptera: Fulgoromorpha: Dictyopharidae)
Leonidas-Romanos Davranoglou, Igor Malenovský, Zestin Wen Wen Soh, Christian M. Schlepütz, and Michael R. Wilson (doi:-)
Abstract丨PDF(-MB)(Proof production in process)
Ground-dwelling Spider Communities Respond to Changes in Riparian Vegetation Widths
Mateus Marques Pires, Everton Nei Lopes Rodrigues, Matheus dos Santos Rocha, José Alessandro Radici Costi, Gustavo Viegas, Cristina Stenert, and Leonardo Maltchik (doi:-)
AbstractEarly view (991.7KB)
New Genus and Two New Species of Semiterrestrial Crabs (Decapoda: Brachyura: Gecarcinucidae) from the Southern Western Ghats, India
Smrithy Raj, Appukuttannair Biju Kumar, and Peter K. L. Ng (doi:-)
AbstractWe follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version.
Population Genetic Structures and Demographic Expansion of Exotic Jellyfish Carybdea brevipedalia in Korean Coasts Inferred from Mitochondrial COI Analysis 
Buhari Lawan Muhammad, Yoseph Seo, Jinho Chae, and Jang-Seu Ki (doi:-)
AbstractEarly view (841KB)
Use of Habitat Suitability Model as a Tool to Highlight Best Conservation Area for the Red-Bellied Monkey (Cercopithecus erythrogaster erythrogaster) in Southern-Benin
Omobayo Ghislain Zoffoun, Côme Agossa Linsoussi, Georges Nobimè, and Brice Augustin Sinsin (doi:-)
AbstractEarly view (1.80MB)
Transplantation Tests of Precious Coral Fragments Using Small-sized Artificial Substratum
Tatsuki Koido, Sho Toshino, Fujio Kumon, Shu Nakachi, Noriyoshi Yoshimoto, and Takuma Mezaki (doi:-)
AbstractEarly view (3.14MB)
Three New Species of the Sun Coral Genus Tubastraea (Scleractinia: Dendrophylliidae) from Hong Kong, China
Sam King Fung Yiu and Jian-Wen Qiu (doi:-)
Abstract丨We follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version.
Hidden Diversity in a Narrow Valley: Description of New Endemic Palearctic Rock Lizard Darevskia (Squamata: Lacertidae) Species from Northeastern Turkey
Muammer Kurnaz, Mehmet Kürşat Şahin, and Ali İhsan Eroğlu (doi:-)
AbstractEarly view (5.74MB)
Eat Better to Look Better: The Relationship between Food Availability, UV Brightness of the Major Claw, and Mating Success of the Fiddler Crab Austruca lactea
Kun-Chang Li, Yu-Shiuan Chen, Wei-Chi Lin, and Hui-Chen Lin (doi:-)
AbstractEarly view (629KB)
New Insights into the Life-history Traits of the Night Shrimp Processa hemphilli (Caridea: Processidae) in the Western Atlantic
Natália Bosco Talamonte, Régis Augusto Pescinelli, Abner Carvalho-Batista, and Rogerio Caetano Costa (doi:-)
AbstractEarly view (1.54MB)
Reproduction of the Blue Jack Mackerel, Trachurus picturatus, in Western Portugal: Microscopic Gonad Analysis Reveals Indeterminate Fecundity and Skipped Spawning Patterns
Ana Neves, Vera Sequeira, Ana Rita Vieira, Elisabete Silva, Frederica Silva, Ana Marta Duarte, Susana Mendes, Rui Ganhão, Carlos Assis, Rui Rebelo, Maria Filomena Magalhães, Maria Manuel Gil, and Leonel Serrano Gordo (doi:-)
AbstractEarly view (1.58MB)
Utilisation of Woody Plants by the Cape Porcupine in Mesic Savannas in South Africa
Unathi Masiobi Kraai, Zivanai Tsvuura, Tlou Julius Tjelele, Ntuthuko Raphael Mkhize, and Manqhai Kraai (doi:-)
AbstractEarly view (635KB)
Biogeography and Ecological Differentiation of Pseudasphondylia gall midges (Diptera: Cecidomyiidae) Distributed in Taiwan and Japan, with Description of a New Species P. kiwiphila sp. nov. and the Southernmost Record of P. elaeocarpi
Sheng-Feng Lin, Makoto Tokuda, Gene-Sheng Tung, Lan-Yu Pan, Wanggyu Kim, Junichi Yukawa, and Man-Miao Yang (doi:-)
AbstractWe follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version. (-MB)
Phylogenetic Relationships and Taxonomic Position of the Genus Parahubrechtia (Nemertea, Palaeonemertea) with Descriptions of Two New Species
Alexei V. Chernyshev, Neonila E. Polyakova, and Shi-Chun Sun (doi:-)
AbstractWe follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version. (-MB)
Descriptive Morphology of Male and Female Genitalia in the Tribe Aphractini (Orthoptera: Tettigoniidae), with a Key to the Genera
Alejandro Andrés Vera and Paula Prelo (doi:-)
AbstractEarly view (2.42MB)
Roosting-site Selectionby Overwintering Black-necked Cranes in the Caohai  Wetland, Guizhou Province, China: Implications for Conservation Management
Xue Gou, Yuan Zhu, Xijiao Sun, Chanshi Hu, Mingming Zhang, Romaan Hayat Khattak, and Haijun Su (doi:-)
AbstractEarly view (610KB)
Foraging Strategy of Black-faced Spoonbill During Breeding Period in Rice Fields of Korea
Sung-Yeon Yoo, Hyung-Kyu Nam, Jong-Kyung Hwang, Jeong-Chil Yoo, and In-Ki Kwon (doi:-)
AbstractEarly view (936KB)
Population and Conservation Status of Pteropus dasymallus in Taiwan
Hui-Wen Wu, Dau-Jye Lu, Ching-Lung Lin, Hsi-Chi Cheng, Chung-Hao Juan, Tsung-Jen Shen, Hua-Ching Lin, and Shiang-Fan Chen (doi:-)
AbstractEarly view (633KB)
The Impact of Lantana camara on Invertebrates and Plant Species of the Groenkloof Nature Reserve, South Africa
Tlou Daisy Raphela and Kevin Duffy (doi:-)
AbstractEarly view (4.59MB)
Host-parasite Interactions Between a Copepod (Pharodes tortugensis) and Small Reef-associated Gobies (Coryphopterus) in the British Virgin Islands
Graham E. Forrester and Rachel J. Finley (doi:-)
AbstractEarly view (2.85MB)
On the Genus Lewisister (Coleoptera, Histeridae), with Description of a New Species from Taiwan
Yu-Hsiang Ho and Masahiro Ôhara (doi:-)
AbstractWe follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version. (-MB)
Three New Polychaete Species of Platynereis (Annelida, Polychaeta, Nereididae) from Taiwan
Pan-Wen Hsueh (doi:-)
AbstractWe follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version. (-MB)
First Report of the Tramp ant Technomyrmex vitiensis Mann, 1921 (Formicidae: Dolichoderinae) in Brazil with Cytogenetic and Sperm Structure Data and an Updated Key to Brazilian Dolichoderinae Genera
Luísa Antônia Campos Barros, Júlio Cezar Mário Chaul, Gisele Amaro Teixeira, Rodrigo Batista Lod, Jérôme Orivel, and Hilton Jeferson Alves Cardoso de Aguiar (doi:-)
AbstractEarly view (1.44MB)
Biogeographic Inferences on the Evolutionary History of the King Cobra (Ophiophagus hannah, Cantor 1836) Species Complex
Fèlix Amat and Daniel Escoriza (doi:-)
AbstractEarly view (3.53MB)
A New Species of Elpidium (Crustacea: Ostracoda: Limnocytheridae) from Brazil and a Morphological Phylogenetic Proposal for the Genus
Julia S. Pereira, Carlos E. F. Rocha, Ricardo L. Pinto, and Marcio B. DASilva (doi:-)
AbstractWe follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version. (-MB)
Searching Behavior in the Tropical Fire Ant Solenopsis geminata (Hymenoptera: Formicidae)
Li‑Chuan Lai, Tzu-Yen Chao, and Ming-Chung Chiu (doi:10.6620/ZS.2022.61-26)
AbstractPDF (6.43MB)
Elevational Distribution of Amphibians: Resolving Distributions, Patterns, and Species  Communities in the Republic of Korea
Desiree Andersen, Ming-Feng Chuang, Minjee Choe, Ajoung Kim, Sera Kwon, Yikweon Jang, and Amaël Borzée (doi:10.6620/ZS.2022.61-25)
AbstractPDF (1.62MB)
Two New Pseudoscorpion Species of the Coastal Genus Garypus L. Koch, 1873 (Garypidae) and an Updated Checklist of the Pseudoscorpiones of Taiwan
Hsiang-Yun Lin, Jun-Xuan Huang, Hsi-Hsuan Liu, and Chih-Han Chang (doi:10.6620/ZS.2022.61-24)
AbstractPDF (6.55MB)
Seasonal Distribution of the Broad-Tailed Hummingbird (Selasphorus platycercus): A Climatic Approach
Norma Hernández-Hernández, Claudio Mota-Vargas, Carlos Lara, and Octavio Rojas-Soto (doi:10.6620/ZS.2022.61-23)
AbstractPDF (3.75MB)
Macrobiotus naginae sp. nov., a New Xerophilous Tardigrade Species from Rokua Sand Dunes (Finland)
Matteo Vecchi, Daniel Stec, Vuori Tommi, Serge Ryndov, Justine Chartrain, and Sara Calhim (doi:10.6620/ZS.2022.61-22)
AbstractPDF (20.2MB)
A New Branchipolynoe (Aphroditiformia: Polynoidae) Scale Worm from the Onnuri Deep-sea Hydrothermal Vent Field, Northern Central Indian Ridge
Sang Lyeol Kim, Hyeongwoo Choi, Seong-il Eyun, Dongsung Kim, and Ok Hwan Yu (doi:10.6620/ZS.2022.61-21)
AbstractPDF (4.60MB)
A New Species of Free-living Marine Nematode (Ptycholaimellus: Chromadoridae: Chromadorida: Nematoda) from Mangrove Wetlands in China
Yi-Jia Shih, Yu-Zhen Chen, and Yu-Qing Guo (doi:10.6620/ZS.2022.61-20)
AbstractPDF (1.45MB)
Bark-stripping Behavior of Formosan Sambar (Rusa unicolor swinhoii) at Tataka, Yushan National Park in Taiwan
Guo-Jing Weng, Shu-Mei Chen, Li-Min Yin, I-Chen Wu, and Ting-An Chou (doi:10.6620/ZS.2022.61-19)
AbstractPDF (2.45MB)
Anatomy and Histology of the Eye of the Nutria Myocastor coypus: Evidence of an Adaptation to a Semi-aquatic Life
Taeko Miyazaki, Yukako Naritsuka, Michihiro Yagami, Shuji Kobayashi, and Koichi Kawamura (doi:10.6620/ZS.2022.61-18)
AbstractPDF (4.99MB)
Changes in Red Color Sensitivity over the Spawning Cycle of Female Three-spined Stickleback (Gasterosteus aculeatus)
Li-Chun Li, Chrysoula Roufidou, Bertil Borg, and Yi Ta Shao (doi:10.6620/ZS.2022.61-17)
AbstractPDF (1.57MB)
Correlated Expression of the Opsin Retrogene LWS-R and its Host Gene in Two Poeciliid Fishes
Chia-Hao Chang (doi:10.6620/ZS.2022.61-16)
AbstractPDF (1.88MB)
A New Sponge Associated Shrimp of the Genus Periclimenaeus Borradaile, 1915 (Decapoda, Palaemonidae) from the Tropical Eastern Atlantic, Cabo Verde Islands
Keider Neves (doi:10.6620/ZS.2022.61-15)
AbstractPDF (9.67MB)
Different Dynamics of Reproductive Cell Development in Oviparous Clausilia bidentata and Ovoviviparous Ruthenica filograna Snails
Krystyna Szybiak, Elżbieta Gabała, and Zbigniew Adamski (doi:10.6620/ZS.2022.61-14)
AbstractPDF (1.20MB)
Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species
Peter K. L. Ng and Jongkar Grinang (doi:10.6620/ZS.2022.61-13)
AbstractPDF (55.6MB)
Phylogenetic Appraisal of Lysiosquillidae Giesbrecht, 1910, and A New Species of Lysiosquilloides Manning, 1977, from Taiwan (Crustacea: Stomatopoda: Lysiosquilloidea)
Shane T. Ahyong and Chia-Wei Lin (doi:10.6620/ZS.2022.61-12)
AbstractPDF (3.72MB)
Integrative Taxonomy, Distribution and Ontogenetic Colouration Change in Neotropical Mountain Catfishes of the Trichomycterus nigroauratus Group (Siluriformes: Trichomycteridae)
Wilson J. E. M. Costaa, José Leonardo O. Mattosa, Sâmela Lopesa, Pedro F. Amorima, and Axel M. Katza (doi:10.6620/ZS.2022.61-11)
AbstractPDF (8.29MB)
First Skeletal Fossil Record of the Red Seabream Pagrus major (Sparidae, Perciformes) from the Late Pleistocene of Subtropical West Pacific, Southern Taiwan
Chien-Hsiang Lin, Hsin-Yueh Ou, Chia-Yen Lin, and Hong-Ming Chen (doi:10.6620/ZS.2022.61-10)
AbstractPDF (60.2MB)
A New Species of Parascorpaena Bleeker, 1876 (Teleostei: Scorpaenidae) from Taiwan
Tak-Kei Chou and Te-Yu Liao (doi:10.6620/ZS.2022.61-09)
AbstractPDF (13.9MB)
Coexistence of Juvenile with Adult Ocypode gaudichaudii at Culebra Beach, Panama: A Temporal-spatial Partitioning Compromise
Adeline Y.P. Yong and Shirley S.L. Lim (doi:10.6620/ZS.2022.61-08)
AbstractPDF (16.9MB)
Predation of Oysters Using an Autonomic Pharynx in the Oyster Leech Cryptostylochus sp. (Polycladida: Stylochidae)
Ching-Jung Teng, Ying-Jung Su, Chih-Yu Yeh, and Wei-Ban Jie (doi:10.6620/ZS.2022.61-07)
AbstractPDF (5.93MB)
The Identity of Homoioplax haswelli (Miers, 1884) (Crustacea: Decapoda: Brachyura)
Peter K. L. Ng, Paul F. Clark, and Shane T. Ahyong (doi:10.6620/ZS.2022.61-06)
AbstractPDF (13.5MB)
Historical Biogeography of the Leptodactylus fuscus Group (Anura, Leptodactylidae): Identification of Ancestral Areas and Events that Modeled their Distribution
Matías E. Cáffaro, Regina G. Medina, María L. Ponssa, and Juan M. Díaz Gómez (doi:10.6620/ZS.2022.61-05)
AbstractPDF (4.26MB)
Genetic Structure of the Mangrove Killifish Kryptolebias hermaphroditus Costa, 2011 (Cyprinodontiformes: Aplocheiloidei) Supports A Wide Connection among its Populations
Pedro F. Amorim, Axel M. Katz, Felipe P. Ottoni, and Pedro H.N. de Bragança (doi:10.6620/ZS.2022.61-04)
AbstractPDF (6.27MB)
Effect of Blue Light on the Growth of the Red Swamp Crayfish Procambraus clarkii Larvae - Seasonal and Sexual Differences
Kenji Toyota, Kazuki Usami, Kanta Mizusawa, and Tsuyoshi Ohira (doi:10.6620/ZS.2022.61-03)
AbstractPDF (1.38MB)
Hiding in Plain Sight: Rain Water Puddles in Nicobar Islands of India Reveal Abundance of a New Frog Species of the Genus Microhyla Tschudi, 1838 (Anura: Microhylidae)
Sonali Garg, Chandrakasan Sivaperuman, G. Gokulakrishnan, S. R. Chandramouli, and S. D. Biju (doi:10.6620/ZS.2022.61-02)
AbstractPDF (71.8MB)
Populational Evidences Support Monogamy Mating System in Five Species of Snapping Shrimps of the Genus Alpheus (Caridea: Alpheidae)
Ana C. Costa-Souza, José R.B. Souza, and Alexandre O. Almeida (doi:10.6620/ZS.2022.61-01)
AbstractPDF (17.8MB)