We publish high quality peer-reviewed articles on animal behavior, comparative physiology, evolution, ecology, and systematics and biogeography on animals taxa from all parts of the world.
OPEN ACCESS (Free Page Charge)
Zoological Studies is an OPEN ACCESS international journal and FREE OF PAGE CHARGES from authors.

Edited by:
Benny K.K. Chan (Editor-in-Chief)
Sen-Lin Tang (Co-Editor-in-Chief)

Impact Factor: 1.6
ISI Journal Citation Reports
Ranking: 2022: 62/176 (Zoology)

ISSN (E): 1810-522X
Follow us on Twitter @ZooStudies and Facebook

Zoological Studies is listed in PubMed
Sumbit manuscript

Zoological Studies Editorial Team

The Editor in Chief, Dr. Benny K.K. Chan, and the Co-editors in Chief, Dr. Sen-Lin Tang are professors/ research fellows at Biodiversity Research Center, Academia Sinica, Taiwan.

The editors and co-editors in chief are supported by 30 Associate Editors including Drs.Sheng-Feng Shen, I-Min Tso, Pung-Pung Hwang, Yeong-Choy Kam, David Miller, Shou-Hsien Li, Chung-Pin Lin, John Wang, Claudia Castellani, Yasuyuki Hashiguchi, Chih-Ming Hung, Chiun-Cheng Ko, Chi-Chien Kuo, Teng-Chiu Lin, Jian-Nan Liu, Ryuji Machida, Hiroomi Miyamoto, Yoko Nozawa, Gray A. Williams, Benny K.K. Chan, Allen Chen, Ka Hou Chu, Jens Thorvald Høeg, Jen-Pan Huang, Hin-Kiu Mok, Felipe Polivanov Ottoni, James D. Reimer, Shen-Horn Yen, Sen-Lin Tang, and Daryi Wang.

(See Editorial Board for details)

Message from the Editor in Chief

We take up a responsibility for the academic society

Nowadays, journals often require expensive page charges/open-access fees to publish scientific research articles. Is it possible there are quality assured, peer-reviewed and open-access SCI scientific journals, in which colleagues can publish their research without these hefty fees? We, the Biodiversity Research Center, Academia Sinica, Taiwan, have taken up this responsibility. We publish a journal entitled Zoological Studies, to provide a free SCI open-access publication platform for colleagues to share their research achievements.
    Publishing articles free of page charges does not mean that articles in Zoological Studies lack quality. All accepted articles have passed a formal review system. Impact factors are often used to judge the quality of a journal. However, article citations can be also affected by the  marketing methods used in journals published by commercial publishers. We expect to maintain a respectable impact factor for Zoological Studies. All articles are 1) formally assigned a DOI number and are open-access on the web, 2) listed and housed in PubMed database and 3) promoted in our twitter and facebook accounts. We are not under the umbrella of any commercial publisher. All the editors and reviewers are volunteers. We hope you can support us by submitting your quality research to Zoological Studies, and subsequently upload your published articles in Research Gate to enhance citations as well.

Prof. Benny K.K. Chan

Recent published articles in 2023
Revision of Salmoneus cristatus (Coutière, 1897) and Allied Forms (Decapoda: Caridea: Alpheidae)
Hossein Ashrafi, Arthur Anker, and Zdeněk Ďuriš (doi:-)
AbstractWe follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version.
Morphological and Molecular Evidence for the Identity of Two Land Hermit Crabs Coenobita longitarsis De Man, 1902 and C. pseudorugosus Nakasone, 1988 (Crustacea: Decapoda: Anomura: Coenobitidae)
Hsi-Te Shih, Kai Chang, Félix Adhi Pramono, and Maria Celia (Machel) D. Malay (doi:-)
AbstractEarly view (2.95MB)
Morphological Description and Molecular Characterisation of Glyptothoa gen. nov., a Fish Parasitic Deep-sea Cymothoid (Crustacea: Isopoda) from the Indian Ocean, with four species, including one new species
Ameri Kottarathil Helna, Panakkool Thamban Aneesh, Appukuttannair Biju Kumar, and Susumu Ohtsuka (doi:-)
AbstractWe follow the code of the International Commission on Zoological Nomenclature, taxonomic papers with new species/genus descriptions will not have early view version.
Update of the Loggerhead Sea Turtle (Caretta caretta) Population Nesting in Koroni, Greece, Mediterranean
Dimitris Margaritoulis, Gonçalo Lourenço, and ALan F. Rees (doi:-)
AbstractEarly view (2.93MB)
Integrative Systematics and Biogeography of the Hydrozoans (Leptothecata: Eirenidae) Eirene menoni Kramp, 1953 and Eirene lacteoides Kubota and Horita, 1992 from Japan and China with Comments on Pacific Ocean Distributions
Gerald L. Crow, Brenden S. Holland, Gaku Yamamoto, Shuhei Ikeda, Aya Adachi, and Kelley Niide (doi:-)
AbstractEarly view (928KB)
Mitophylogeny of Pangasiid Catfishes and its Taxonomic Implications for Pangasiidae and the Suborder Siluroidei
Thuy Yen Duong, Linh Thi Khanh Pham, Xuyen Thi Kim Le, Ngoc Tran Thi Nguyen, Siti Azizah Mohd Nor, and Thanh Hoa Le (doi:-10.6620/ZS.2023.62-48)
AbstractPDF (4.53MB)
The Effect of Light Pollution on the Sea Finding Behavior of Green Turtle Hatchlings on Lanyu Island, Taiwan
Chih-Hao Yen, Yin-Ting Chan, Yao-Chi Peng, Kuo-Hui Chang, and I-Jiunn Cheng (doi:10.6620/ZS.2023.62-47)
AbstractPDF (2.75MB)
Migration Behavior of Anguilla celebesensis Silver Eels within their Tomini Bay Spawning Area
Ryotaro Manabe, Takatoshi Higuchi, Shun Watanabe, Fadly Y. Tantu, Hagi Y. Sugeha, Hiroki Kaneko, Michael J. Miller, Seishi Hagihara, Tatsuki Yoshinaga, Augy Syahailatua, Sam Wouthuyzen, Triyanto, Kawilarang W. A. Masengi, Katsufumi Sato, Jun Aoyama, and Katsumi Tsukamoto (doi:10.6620/ZS.2023.62-46)
AbstractPDF (44.6MB)
Minuca panema (Coelho, 1972): Resurrection of a Fiddler Crab Species from Brazil Closely Related to Minuca burgersi (Holthuis, 1967) (Crustacea, Decapoda, Brachyura, Ocypodidae)
Carl. L. Thurman, Hsi-Te Shih, and John C. McNamara (doi:10.6620/ZS.2023.62-45)
AbstractPDF (24.1MB)
Drainage Network Morphology Influences Population Structure and Gene Flow of the Andean Water Frog Telmatobius pefauri (Anura: Telmatobiidae) of the Atacama Desert, Northern Chile
Pablo Fibla, Paola A. Sáez, Franco Cruz-Jofré, and Marco A. Méndez (doi:10.6620/ZS.2023.62-44)
AbstractPDF (13.2MB)
Geographic Distribution and Genetic Structures of the Tideland Snails Pirenella nipponica and P. asiatica in Taiwan and Japan
Shintaro Kato, Mizuki Ohta, Hiroaki Fukumori, Te-Hua Hsu, Tin-Yam Chan, and Shigeaki Kojima (doi:10.6620/ZS.2023.62-43)
AbstractPDF (2.37MB)
Chromosomal Study on Selected Small Araneomorph Spiders from Brazil, Including the First Records in Palpimanidae and Theridiosomatidae (Araneae, Araneomorphae)
Débora Duarte Dutra, Antônio Domingos Brescovit, and Douglas Araujo (doi:10.6620/ZS.2023.62-42)
AbstractPDF (3.18MB)
Organismal Responses to Coastal Acidification Informed by Interrelating Erosion, Roundness and Growth of Gastropod Shells
David J. Marshall and Amira Rashid (doi:10.6620/ZS.2023.62-41)
AbstractPDF (3.07MB)
Integrative Taxonomy of Cyclocyprididae Kaufmann, 1900 (Ostracoda: Podocopa) with Description of a New Genus and Species
Maria Bisquert-Ribes, Juan Rueda, Ferran Palero, Sukonthip Savatenalinton, and Francesc Mesquita-Joanes (doi:10.6620/ZS.2023.62-40)
AbstractPDF (11.9MB)
Phylogeography of Long-spined Sea Urchin Diadema setosum Across the Indo-Malay Archipelago
Indra Bayu Vimono, Philippe Borsa, Régis Hocdé, and Laurent Pouyaud (doi:10.6620/ZS.2023.62-39)
AbstractPDF (2.04MB)
Cellular and Biochemical Changes in Early Embryonic Development of a Scleractinian Coral, Fimbriaphyllia (Euphyllia) ancora
Shinya Shikina, Yi-Ling Chiu, Yi-Jou Chung, Yi-ChenYao, Tai-Yu Liu, Yan-Horn Lee, Masatoshi Mita, and Ching-Fong Chang (doi:10.6620/ZS.2023.62-38)
AbstractPDF (4.74MB)
Taxonomy and Phylogeography of the Freshwater Crab Geothelphusa tawu Species Complex (Crustacea: Decapoda: Potamidae) from Southern Taiwan and Offshore Islets
Hsi-Te Shih, Jhih-Wei Hsu, Kai Chang, and Min-Wan Chen (10.6620/ZS.2023.62-37)
AbstractPDF (8.11MB)
Brakemyia, a New Neotropical Jackal Fly Genus of Milichiidae (Insecta: Diptera) Associated with Carton Ant Nest
Fernando S. Carvalho-Filho, Rodrigo R. Barbosa, and Matheus M.M. Soares (doi:10.6620/ZS.2023.62-36)
AbstractPDF (6.69MB)
Deuterogyny and the Association of Two Vagrant Eriophyoid Mites (Acariformes, Eriophyoidea) with the Host-plant Generative Organs of Two Broad-leaved Trees in North-West Russia
Philipp E. Chetverikov, Alexey G. Desnitskiy, Pavel B. Klimov, Sebahat K. Ozman-Sullivan, Anna E. Romanovich, and Sogdiana I. Sukhareva (doi:10.6620/ZS.2023.62-35)
AbstractPDF (9.6MB)
Multigene Phylogenies of the Estuarine Sesarmid Parasesarma bidens Species Complex (Decapoda: Brachyura: Sesarmidae), with Description of Three New Species
Hsi-Te Shih, Jhih-Wei Hsu, and Jheng-Jhang Li (doi:10.6620/ZS.2023.62-34)
AbstractPDF (102MB)
Identification of Phyllosoma Larvae of the Lobsters Acantharctus and Biarctus (Crustacea: Scyllaridae) from the SW Indian Ocean
Rebeca Genis-Armero, Johan Groeneveld, Sohana Singh, Paul F. Clark, Laure Corbari, and Ferran Palero (doi:10.6620/ZS.2023.62-33)
AbstractPDF (6.28MB)
Phylogeography and Genetic Structure of the Bush Cricket Decma fissa (Orthoptera, Tettigoniidae) in Southern China
Qi Guo, Qi-Di Zhu, Zhi-Jun Zhou, and Fu-Ming Shi (doi:10.6620/ZS.2023.62-32)
AbstractPDF (15MB)
Taxonomic Revision, Morphology and Natural History of the Stick Insect Genus Xerosoma Serville, 1831 (Insecta: Phasmatodea)
Phillip Watzke Engelking, Victor Morais Ghirotto, Edgar Blois Crispino, Thies H. Büscher, Raphael Aquino Heleodoro, Pedro Alvaro Barbosa Aguiar Neves, and Pitágoras da Conceição Bispo (doi:10.6620/ZS.2023.62-31)
AbstractPDF (55.7MB)
Assessing the Taxonomic Status of the Gray Brocket Mazama simplicicornis argentina Lönnberg, 1919 (Artiodactyla: Cervidae)
Eluzai Dinai Pinto Sandoval, Gabrielle Queiroz Vacari, Juan Pablo Juliá, Susana González, Miluse Vozdova, Halina Cernohorska, Svatava Kubickova, Daniela C. Kalthoff, and José Mauricio Barbanti Duarte (doi:10.6620/ZS.2023.62-30)
AbstractPDF (3.46MB)
Taxonomy and Molecular Phylogeny of the Sea Anemone Macrodactyla (Haddon, 1898) (Cnidaria, Actiniaria), with a Description of a New Species from Singapore
Nicholas Wei Liang Yap, Michela Lee Mitchell, Zheng Bin Randolph Quek, Ria Tan, Koh Siang Tan, and Danwei Huang (doi:10.6620/ZS.2023.62-29)
AbstractPDF (26.5MB)
Effects of Different LED Light Spectra on Growth and Immunity of the Japanese Eel (Anguilla japonica) and Giant Mottled Eel (A. marmorata)
Yen-Ting Lin, Wei-Chun Hung, Yin-Fu Yeh, Kuang-Mao Lu, Ding-Hwa Cherng, and Yu-San Han (doi:10.6620/ZS.2023.62-28)
AbstractPDF (1.85MB)
A New Distinctive Darwin Wasp Represents the First Record of the Ophion minutus Species-group (Hymenoptera: Ichneumonidae: Ophioninae) from Japan and the Far East, with an Analysis of DNA Barcode-based Species Delimitation in Ophion
So Shimizu and Kaoru Maeto (doi:10.6620/ZS.2023.62-27)
AbstractPDF (87.3MB)
A New Paralepetopsis Limpet from a South China Sea Seep Hints at a Paraphyletic Neolepetopsidae
Chong Chen, Zhaoyan Zhong, Jian-Wen Qiu, and Jin Sun (doi:10.6620/ZS.2023.62-26)
AbstractPDF (10.7MB)
Long-Read Genome Sequencing of Abscondita cerata (Coleoptera: Lampyridae), the Endemic Firefly of Taiwan
Tzi-Yuan Wang, King-Siang Goh, Liang-Jong Wang, Li-Ling Wu, Feng-Yu Wang, and Yu-Wei Wu (doi:10.6620/ZS.2023.62-25)
AbstractPDF (3.44MB)
Genetic Structure of the Endemic Fiddler Crab Uca (Xeruca) formosensis on the West Coast of Taiwan
Kun-Chin Hung, Ching-Yu Liou, Chih-Chung Wen, and Hui-Chen Lin (doi:10.6620/ZS.2023.62-24)
AbstractPDF (2.92MB)
Discovery of a New Sea Spider Belonging to the Genus Pycnogonum (Pycnogonida: Pycnogonidae) in the Mesophotic Zone of Jejudo Island, Korea
Damin Lee, Jin-Ho Park, and Taeseo Park (doi:10.6620/ZS.2023.62-23)
AbstractPDF (4.5MB)
Seaward Migration and Larval Release Coincide with Lunar and Light-dark Cycles in Supratidal Land Crabs Cardisoma carnifex and Epigrapsus notatus
Chung-Chieh Chang, Kent A. Hatch, Chia-Hsuan Hsu, Wenbe Hwang, Hung-Chang Liu, and Yuan-Mou Chang (doi:-10.6620/ZS.2023.62-22)
AbstractPDF (6.19MB)
Molecular and Morphological Characterizations of the Fish Parasitic Isopod Mothocya parvostis (Crustacea: Cymothoidae) Parasitizing Optional Intermediate Hosts: Juveniles of the Cobaltcap Silverside Hypoatherina tsurugae and Yellowfin Seabream Acanthopagrus latus
Hiroki Fujita, Kentaro Kawai, Diego Deville, and Tetsuya Umino (doi:10.6620/ZS.2023.62-21)
AbstractPDF (27.9MB)
Revision of the Genus Lacunipotamon Dai, Song, He, Cao, Xu & Zhong, 1975 (Crustacea: Decapoda: Brachyura: Potamidae), with Descriptions of Eight New Species from Vietnam
Peter K. L. Ng, Zhi Wan, Tan, and Van Tri Ngo (doi:10.6620/ZS.2023.62-20)
AbstractPDF (102.1MB)
Past, Present and Future of the Aral Sea - A Review of its Fauna and Flora before and during the Regression Crisis
Igor S. Plotnikov, Nikolai V. Aladin, Lubov V. Zhakova, Jens Mossin, and Jens T. Høeg (doi:10.6620/ZS.2023.62-19)
AbstractPDF (12.4MB)
Understanding the Symbiotic Relationship between the Sea Urchin Loxechinus albus (Molina, 1782) and the Pea Crab Pinnaxodes chilensis (H. Milne Edwards, 1837): a Potential Parasitism
Hans N. Jaramillo, Luis P. Salas-Yanquin, Joseline A. Büchner-Miranda, Víctor M. Cubillos, Jaime A. Montory, Jan A. Pechenik, Luis M. Pardo, and Oscar R. Chaparro (doi:10.6620/ZS.2023.62-18)
AbstractPDF (3.53MB)
A New Species of Box Jellyfish (Cnidaria: Tripedaliidae: Tripedalia) from Hong Kong, China
Yanan Sun, Justin Hon Yin Tsui, Rachel Ting Huen Wong, Ringo Nga Ching Cheung, Murphy Kam Pui Ng, Carmen K.M. Or, and Jian-Wen Qiu (doi:10.6620/ZS.2023.62-17)
AbstractPDF (5.45MB)
Adding the Molecular Diversity Information of the Common Fouling Barnacle Amphibalanus amphitrite (Darwin, 1854) (Crustacea: Cirripedia) from the Persian Gulf and Gulf of Oman to the Global Diversity Pattern
Asma Moeinadini, Alireza Sari, Adnan Shahdadi, Ahmad-Reza Katouzian, Alimorad Sarafrazi, and Elahe Elahi (doi:10.6620/ZS.2023.62-16)
AbstractPDF (4.14MB)
A New Polyclad Flatworm, Idiostylochus tortuosus gen. nov., sp. nov. (Platyhelminthes, Polycladida) from France. Can this Foreign Flatworm be Responsible for the Deterioration of Oyster and Mussel Farms?
Adrian Gutiérrez, Isabelle Auby, Benoit Gouillieux, Guillemine Daffe, Cecile Massé, Elvire Antajan, and Carolina Noreña (doi:10.6620/ZS.2023.62-15)
AbstractPDF (5.32MB)
Cryptic Genetic Diversity in the Coastal Isopod Alloniscus oahuensis from the Pacific Ocean
Carlos A. Santamaria and Maddy R. Koch (doi:10.6620/ZS.2023.62-14)
AbstractPDF (1.51MB)
Genetic Assessment of Fertile F1 Hybrids between Two Hercules Beetles, Dynastes maya Hardy and D. grantii Horn (Scarabaeidae)
Jen-Pan Huang, Wei-Yun Chen, and My Hanh Le (doi:10.6620/ZS.2023.62-13)
AbstractPDF (115MB)
Low Diversity of Major Histocompatibility Complex (MHC) Genes in Endangered Malayan Tapir (Tapirus indicus)
Nurul Adilah Ismail, Christina Seok Yien Yong, Simon Yung Wa Sin, and Geetha Annavi (doi:10.6620/ZS.2023.62-12)
AbstractPDF (2.14MB)
Genetic Profile of the Parasitic Varroan Mite Varroa destructor (Arachnida: Mesostigmata: Varroidae) in Taiwan: a New Taiwanese Haplotype Intermediate Between the Highly Virulent Russian and Less Virulent Japanese Types Identified in the Honey Bee Host Apis cerana
Tsen Hua, Panuwan Chantawannakul, Cheng-Lung Tsai, and Wen-Bin Yeh (doi:10.6620/ZS.2023.62-11)
AbstractPDF (5.18MB)
Tuerkayana latens, a New Species of Land Crab from French Polynesia, with a Discussion on the Phylogeny of the Genus (Crustacea: Decapoda: Brachyura: Gecarcinidae)
Peter K. L. Ng and Hsi-Te Shih (doi:10.6620/ZS.2023.62-10)
AbstractPDF (25.3MB)
Composition and Dynamics of Hexapod Communities on Yushan Bamboo (Yushania niitakayamensis) in the Subtropical Montane Areas of Taiwan
Tsen Hua, Hui-Yi Li, Cheng-Lung Tsai, Wing Wei Wang, Chao-Yuan Lin, and Wen-Bin Yeh (doi:10.6620/ZS.2023.62-09)
AbstractPDF (8.52MB)
Terelabrus toretore sp. nov. (Perciformes: Labridae), a New Species of Striped Hogfish from Tahiti, with Range Extensions for Two Congeners
Bart Shepherd, Hudson T. Pinheiro, Tyler A. Y. Phelps, Gilles Siu, and Luiz A. Rocha (doi:10.6620/ZS.2023.62-08)
AbstractPDF (7.13MB)
Lanternflies (Hemiptera: Fulgoridae) of Taiwan
You-Sheng Lin, Jhih-Rong Liao, Shiuh-Feng Shiao, and Chiun-Cheng Ko (doi:10.6620/ZS.2023.62-07)
AbstractPDF (100MB)
Annotated Catalog of Vespid Hosts (Hymenoptera: Vespidae) of Tachinidae (Diptera), with Description of a New Species of Ophirion Townsend from Brazil
Filipe Macedo Gudin (doi:10.6620/ZS.2023.62-06)
AbstractPDF (7.78MB)
Using the Integrative Approach to Update a Gap of One Century: Redescription and New Distribution Records of the South American Tarantulas Grammostola pulchra (Araneae: Mygalomorphae: Theraphosidae)
Renan S. Pittella, Pedro G. Bassa, Edison Zefa, and Filipe M. Bianchi (doi:10.6620/ZS.2023.62-05)
AbstractPDF (9.77MB)
Effect of pH on the Early Development of the Biofouling Ascidian Ciona robusta
Bailey S.C.L. Jones, Lauren A. Holt, and Kit Yu Karen Chan (doi:10.6620/ZS.2023.62-04)
AbstractPDF (1.12MB)
First Satellite Tagging of the Northern Fur Seals (Callorhinus ursinus) on the Tyuleniy Island, the Sea of Okhotsk
Peter A. Permyakov, Sergey D. Ryazanov, Alexey M. Trukhin, Vyacheslav B. Lobanov, Hyun Woo Kim, and Seok-Gwan Choi (doi:10.6620/ZS.2023.62-03)
AbstractPDF (7.93MB)
Identification of Sexually Dimorphic Genes in Pectoral Fin as Molecular Markers for Assessing the Sex of Japanese Silver Eels (Anguilla japonica)
Hsiang-Yi Hsu, Chia-Hsien Chuang, I-Hsuan Lu, Chung-Yen Lin, and Yu-San Han (doi:10.6620/ZS.2023.62-02)
AbstractPDF (5.59MB)
Twenty Years of Sea Turtle Strandings in New Caledonia
Tyffen Read, Richard Farman, Jean-Christophe Vivier, Frederic Avril, Hugues Gossuin, and Laurent Wantiez (doi:10.6620/ZS.2023.62-01)