We publish high quality peer-reviewed articles on animal behavior, comparative physiology, evolution, ecology, and systematics and biogeography on animals taxa from all parts of the world.
OPEN ACCESS (Free Page Charge)
Zoological Studies is an OPEN ACCESS international journal and FREE OF PAGE CHARGES from authors.

Edited by:
Benny K.K. Chan (Editor-in-Chief)
Sen-Lin Tang (Co-Editor-in-Chief)

Impact Factor: 2.058
ISI Journal Citation Reports
Ranking: 2020: 53/175 (Zoology)

ISSN (E): 1810-522X
Follow us on Twitter @ZooStudies and Facebook

Zoological Studies is listed in PubMed
Sumbit manuscript

Zoological Studies Editorial Team

The Editor in Chief, Dr. Benny K.K. Chan, and the Co-editors in Chief, Dr. Sen-Lin Tang are professors/ research fellows at Biodiversity Research Center, Academia Sinica, Taiwan.

The editors and co-editors in chief are supported by 30 Associate Editors including Drs.Sheng-Feng Shen, I-Min Tso, Pung-Pung Hwang, Yeong-Choy Kam, David Miller, Shou-Hsien Li, Chung-Pin Lin, John Wang, Claudia Castellani, Yasuyuki Hashiguchi, Chih-Ming Hung, Chiun-Cheng Ko, Chi-Chien Kuo, Teng-Chiu Lin, Jian-Nan Liu, Ryuji Machida, Hiroomi Miyamoto, Yoko Nozawa, Gray A. Williams, Benny K.K. Chan, Allen Chen, Ka Hou Chu, Jens Thorvald Høeg, Jen-Pan Huang, Hin-Kiu Mok, Felipe Polivanov Ottoni, James D. Reimer, Shen-Horn Yen, Sen-Lin Tang, and Daryi Wang.

(See Editorial Board for details)

Message from the Editor in Chief

We take up a responsibility for the academic society

    Nowadays, we often need to pay expensive page charges/open-access fees to publish our works as most of the journals are now run by business publishers. Is it possible there are quality assured, peer-reviewed and open-access SCI scientific journals, in which colleagues can publish their research achievement free? We, Biodiversity Research Center, Academia Sinica, Taiwan, attempts to take up this responsibility. We run Zoological Studies, to provide a free SCI open-access publication platform for colleagues to share out their research achievements.
    Publish articles free of page charges does not mean Zoological Studies is lack of good quality. All the accepted articles passed a formal review system. Impact factors is an indication for the article citations in a journals and has implications for journal quality. However, article citations can be also affected by the ways of marketing methods for journals in commercial publishers. We are trying hard to maintain a good impact factor for Zoological Studies. All articles are 1) formally applied for a DOI number and being open-access in the web, 2) listed and housed in PubMed database and 3) promoted in our twitters and facebook. We are not under any umbrella of commercial publishers. All the editors and reviewers are volunteers for the academics. We hope you can support us to submit your quality works to Zoological Studies, and upload your published articles in Research Gate to enhance citations as well.

Prof. Benny K.K. Chan

Recent published articles in 2021
A Soil Nematode Community Response to Reclamation of Salinized Abandoned Farmland
Lei Yang, Fenghua Zhang, and Yanqin Luo (doi:
Abstract丨PDF(-MB)(Proof production in process)
Quantifying the Effect of Land-cover Change on the Endangered Farmland Green Treefrog (Zhangixalus arvalis) in an Agricultural Landscape: Implications for Conservation
Sin Chen, Meng-Hsien Chuang, Hau-Jie Shiu, and Jian-Nan Liu (doi:
Abstract丨PDF(-MB)(Proof production in process)
High Mountain Echiniscid (Heterotardigrada) Fauna of Taiwan
Piotr Gąsiorek, Katarzyna Vončina, Reinhardt Møbjerg Kristensen, and Łukasz Michalczyk (doi:
Abstract丨PDF(-MB)(Proof production in process)
Bacterial Communities in Multiple Tissues Across the Body Surface of Three Coastal Shark Species
Chelsea Black, Liza Merly, and Neil Hammerschlag (doi:
Abstract丨PDF(-MB)(Proof production in process)
A New Species of the Intertidal Limpet Eoacmaea (Patellogastropoda: Eoacmaeidae) from Yonaguni Island, Japan and Taiwan
Tomoyuki Nakano, Takenori Sasaki, Takeharu Kosuge, and Benny K.K. Chan (doi:-)
AbstractEarly view (1.98MB)
Dietary Habits of Free-ranging Banded Langur (Presbytis femoralis) in a Secondary-human Modified Forest in Johor, Malaysia
Mohd Faudzir Najmuddin, Hidayah Haris, Noratiqah Norazlimi, Farhani Ruslin, Ikki Matsuda, Badrul Munir Md-Zain, and Muhammad Abu Bakar Abdul-Latiff (doi:
AbstractEarly view (386KB)
The Hillstream Decapod Crustaceans of Shenzhen, China, with Description of a New Species of Freshwater Crab (Crustacea: Brachyura: Potamidae) of the Genus Megapleonum Huang, Shih & Ahyong, 2018
Chao Huang and Si-Ying Mao (doi:
AbstractEarly view (4.12MB)
A New Species of the Anchovy Genus Stolephorus Lacepède 1803 from North Sumatra, Indonesia, and Redescriptions of Stolephorus pacificus Baldwin 1984 and Stolephorus teguhi Kimura, Hori and Shibukawa 2009 (Teleostei: Clupeiformes: Engraulidae)
Harutaka Hata and Hiroyuki Motomura (doi:
AbstractEarly view (867KB)
Acoustic Properties of the Otolith of the Large Yellow Croaker Larimichthys crocea (Perciformes: Sciaenidae)
Xin-Hai Zhang, Yi Tao, Yang-Liang Zhou, Li-Guo Tang, Min Liu, and Xiao-Mei Xu (doi:
AbstractEarly view (1.64MB)
Population Genetic Structures of the Jellyfish Aurelia coerulea Polyps on Korean Coasts and Implications as Revealed by Mitochondrial COI
Yoseph Seo, Buhari Lawan Muhammad, Jinho Chae, and Jang-Seu Ki (doi:
AbstractEarly view (976KB)
Life History Traits of the Snapping Shrimp Alpheus carlae (Decapoda: Alpheidae) from the South-eastern Coast of Brazil
Yoseph Seo, Buhari Lawan Muhammad, Jinho Chae, and Jang-Seu Ki (doi:
AbstractEarly view (2.08MB)
Histological Evidence of Multiple Spawning in Wild Female Japanese Eel Anguilla japonica
Akio Shimizu, Shigeho Ijiri, Hikari Izumi, Koichiro Gen, Hiroaki Kurogi, Hiroshi Hashimoto, Hideki Tanaka, Tadao Jinbo, Hiroaki Saito, and Seinen Chow (doi:
AbstractEarly view (1.99MB)
Diversity in the Taiwanese Swimming Crabs (Crustacea: Brachyura: Portunidae) Estimated through DNA Barcodes, with Descriptions of 14 New Records
Yu-Hsuan Huang and Hsi-Te Shih (doi:
AbstractEarly view (3.59MB)
A New Species of the Genus Ophiomonas Djakonov (Echinodermata: Ophiuroidea: Amphilepididae) from the Deep-Sea of Japan
Masanori Okanishi, Takumi Matsuo, and Toshihiko Fujita (doi:
AbstractEarly view (2.01MB)
Preliminary Observations on the Effect of Temperature and Food Concentration on the Egg Production Rate and Hatching Success of Acartia amboinensis from the Central Red Sea
Mohsen M. El-Sherbiny and Ali Al-Aidaroos (doi:
AbstractEarly view (894KB)
A New Species of Amphipod Pariphinotus Kunkel, 1910 (Amphipoda: Phliantidae) from Southwestern Atlantic
André R. Senna, Urssula N. Guedes, Luiz F. Andrade, and Guilherme H. Pereira-Filho (doi:
AbstractEarly view (1.09MB)
Effects of Farming Systems on Insect Communities in the Paddy Fields of a Simplified Landscape During a Pest-control Intervention
Chao-Nien Koh, Ming-Chih Chiu, Ling-Mu Jaung, Yung-Jen Lu, and
and Hsing-Juh Lin (doi:10.6620/ZS.2021.60-56)
AbstractPDF (2.14MB)
Diversity and Species Composition of Bark and Ambrosia Beetles Captured Using Ethanol Baited Traps on Different Hosts in East Java, Indonesia
Hagus Tarno, Yogo Setiawan, Cindy Budi Kusuma, Miftachul Fitriyah, Ahmad Hudan Nazaruddin, Alvian Putra Yawandika, Hanif Ardiansyah Nasution, Ronauli Saragih, Achmad Praditya Yoga Bagasta, Zeng Wang, and Jianguo Wang (doi:
AbstractEarly view (803KB)
Sympatric Two-species Infestation by Rhizocephalan Barnacle Parasites in the Spider Crab Pugettia aff. ferox Ohtsuchi & Kawamura, 2019 from Peter the Great Bay (Northwestern Sea of Japan)
Darya D. Golubinskaya, Olga M. Korn, Svetlana N. Sharina, and Nikolai I. Selin (doi:
AbstractEarly view (740KB)
Ecological Role of Cyprideis torosa and Heterocypris salina (Crustacea, Ostracoda) in Saline Rivers of the Lake Elton Basin: Abundance, Biomass, Production, Fatty Acids
Vladimir A Gusakov, Olesia N Makhutova, Michail I Gladyshev, Larisa V Golovatyuk, and Tatiana D Zinchenko (doi:
AbstractEarly view (3.62MB)
Threats to Site Occupation of Carnivores: A Spatiotemporal Encroachment of Non-native Species on the Native Carnivore Community in A Human-dominated Protected Area
Kasereka Vitekere, Luc Mumbere Lango, Jiao Wang, Mengyan Zhu, Guangshun Jiang, and Yan Hua
AbstractPDF (3.16MB)
Influence of Visitors on the Time Budget, Ranging and Strata Use of Lowe’s Monkey (Cercopithecus lowei) at Boabeng-Fiema Monkey Sanctuary, Ghana
Núria Badiella-Giménez, Bright Obeng Kankam, and Llorenç Badiella
AbstractPDF (7.00MB)
Taxonomy, Evolutionary and Dispersal Events of Pig-Tailed Macaque, Macaca nemestrina (Linnaeus, 1766) in Southeast Asia With Description of a New Subspecies, Macaca nemestrina perakensis in Malaysia
Muhammad Abu Bakar Abdul-Latiff and Badrul Munir Md-Zain (doi:
AbstractEarly view (1.83MB)
Reproduction and Early Juvenile Growth of the Giant Clams Tridacna noae and Tridacna maxima in Taiwan
Po-Wei Su, Gwo-Liang Zhang,
Bonien Chen, Keryea Soong, and Li-Lian Liu (doi:10.6620/ZS.2021.60-49)
AbstractPDF (2.10MB)
Ecomorphological Adaptations of Second Maxilliped-setation: Insights from Three Species of Fiddler Crabs from Panama
hirley S.L. Lim and Sarah J.A. Goh (doi:10.6620/ZS.2021.60-48)
AbstractPDF (4.04MB)
Classification and Morphology of Rhinocypha spp. (Odonata): A Comprehensive Taxonomic Study within the Females
oorhidayah Mamat, Arpah-Abu, and Y. Norma-Rashid (doi:10.6620/ZS.2021.60-47)
AbstractPDF (MB)
Moderate Population Structure in Drosophila sturtevanti from the South American Atlantic Forest Biome
runa Memari Trava, Rogério Pincela Mateus, Luciana Paes de Barros Machado, and Lilian Madi-Ravazzi (doi:10.6620/ZS.2021.60-46)
AbstractPDF (3.17MB)
Larval Performance of Amphidromous and Landlocked Atyid Shrimp Species in the Genus Paratya under Different Temperature and Salinity Conditions
atsuyuki Hamasaki, Shuji Kondo, and Shigeki Dan (doi:10.6620/ZS.2021.60-45)
AbstractPDF (7.35MB)
Thermal Physiological Performance and Thermal Metabolic Scope of the Whelk Kelletia kelletii (Forbes, 1850) (Gastropoda: Neptuneidae) Acclimated to Different Temperatures
Fernando Díaz, Ana Denise Re-Araujo, Eugenio Carpizo-Ituarte, Zaul Garcia-Esquivel, Ernesto Larios-Soriano, Leonel Perez-Carrasco, and Ernesto Lerma (doi:10.6620/ZS.2021.60-44)
AbstractPDF (2.09MB)
Phylogenetic Position of Trichomycterus payaya and Examination of Osteological Characters Diagnosing the Neotropical Catfish Genus Ituglanis (Siluriformes: Trichomycteridae)
Wilson J. E. M. Costaa, José Leonardo O. Mattos, and Axel M. Katza (doi:10.6620/ZS.2021.60-43)
AbstractPDF (7.88MB)
From Asia to Europe? Where Could the Geographical Place of Origin of Polymerus (Pachycentrum) carpathicus (Horváth, 1882) (Hemiptera: Miridae) Be?
Agnieszka Bugaj-Nawrocka, Artur Taszakowski, and Grzegorz Gierlasiński (doi:-10.6620/ZS.2021.60-42)
AbstractPDF (37.4MB)
Contact Reactions Between Individuals of the Coral-killing Sponge, Terpios hoshinota
urika Hirose, Siti Nurul Aini, and Hideyuki Yamashiro (doi:10.6620/ZS.2021.60-41)
AbstractPDF (3.23MB)
Identifying A New Phylogeographic Population of the Blyth’s Tragopan (Tragopan blythii) through Multi-locus Analyses
Jingyi Zou, Lu Dong, Geoffrey Davison, Win Hlaing, Moe Myint Aung, Yanyun Zhang, Zhengwang Zhang, Nan Wang, and De Chen (doi:10.6620/ZS.2021.60-40)
AbstractPDF (3.82MB)
Larval Morphology of Pit-building Antlions of the Tribe Myrmeleontini (Neuroptera, Myrmeleontidae) from Taiwan
Yu-Hsiu Lin, Jhih-Rong Liao, and Chiun-Cheng Ko (doi:10.6620/ZS.2021.60-39)
AbstractPDF (25.8MB)
Seasonal Growth of the Purple Sea Urchin Heliocidaris crassispina Revealed by Sequential Fluorochrome Tagging
Juan Diego Urriago Suarez, Jane C.Y. Wong, Gilbert Lui, Clement P. Dumont, Jian-Wen Qiu, and Monthon Ganmanee (doi:
AbstractPDF (12.6MB)
Plasticity of Foraging Strategies Adopted by the Painted Ghost Crab, Ocypode gaudichaudii, in Response to in situ Food Resource Manipulation Experiments
Adeline Y.P. Yong and Shirley S.L. Lim
AbstractPDF (7.98MB)
Otolith Morphology: A Hidden Tool in the Taxonomic Study of Goatfishes (Teleostei: Perciformes: Mullidae)
Sorour Echreshavi, Hamid Reza Esmaeili, Azad Teimori, and Mohsen Safaie (doi:
AbstractPDF (14.8MB)
Review of Chilean Cyrtinae (Diptera: Acroceridae) with the Description of Three New Species and the First Record of Villalus inanis from Argentina
Rodrigo M. Barahona-Segovia, Vicente Valdés Guzmán, and Laura Pañinao-Monsálvez (doi:10.6620/ZS.2021.60-35)
AbstractPDF (46.1MB)
Spatial Distribution and Dietary Niche Breadth of Leopard Cats (Prionailurus bengalensis) Inhabiting Margalla Hills National Park, Pakistan
Hira Fatima, Tariq Mahmood, Lauren Mae Hennelly, Muhammad Farooq, Faraz Akrim, and Muhammad Sajid Nadeem (doi:10.6620/ZS.2021.60-34)
AbstractPDF (3.83MB)
Littoral water in Hong Kong as a Potential Transient Habitat for Juveniles of a Temperate Deepwater Gnomefish, Scombrops boops (Acropomatiformes: Scombropidae)
Jiehong Wei, Jiarui Gu, Min Liu, Bai-An Lin, Gabriel Y. Lee, Tak-Cheung Wai, Paul K.S. Lam, Meng Yan, and Priscilla T.Y. Leung (doi:10.6620/ZS.2021.60-33)
AbstractPDF (2.74MB)
Tissue-specific Isotopic Incorporation Turnover Rates and Trophic Discrimination Factors in the Freshwater Shrimp Macrobrachium borellii (Crustacea: Decapoda: Palaemonidae)
María Florencia Viozzi, Carlos Martínez del Rio, and Verónica Williner (doi:10.6620/ZS.2021.60-32)
AbstractPDF (2.32MB)
Population Genetics Analysis of a Pomacea Snail (Gastropoda: Ampullariidae) in Thailand and its Low Infection by Angiostrongylus cantonensis
Abdulhakam Dumidae, Pichamon Janthu, Chanakan Subkrasae, Raxsina Polseela, Bandid Mangkit, Aunchalee Thanwisai, and Apichat Vitta (doi:10.6620/ZS.2021.60-31)
AbstractPDF (7.70MB)
Species Redescriptions and New Species of the Onthophagus mexicanus Species Group (Coleoptera: Scarabaeidae), with Notes on Distribution and Rodent-dung Beetle Associations
Victor Moctezuma and Gonzalo Halffter (doi:10.6620/ZS.2021.60-30)
AbstractPDF (22.9MB)
Description of a New Species of the Marine Flatworm Prosthiostomum (Platyhelminthes: Polycladida) and its Three Known Congeners from Misaki, Japan, with Inference of Their Phylogenetic Positions within Prosthiostomidae
Aoi Tsuyuki, Hisanori Kohtsuka, and Hiroshi Kajihara (doi:10.6620/ZS.2021.60-29)
AbstractPDF (63.4MB)
Taxonomic Revision of Living Boring Bivalves Belonging to the Family Pholadidae Lamarck, 1809, (Bivalvia: Myida) from the Southwestern Atlantic
Marina Delfino and Javier H. Signorelli (doi:10.6620/ZS.2021.60-28)
AbstractPDF (31.9MB)
Setteleia, a New Genus for Four New Species from the Philippines (Insecta: Lepidoptera: Erebidae: Arctiinae)
Anton V. Volynkin and Karel Černý (doi:10.6620/ZS.2021.60-27)
AbstractPDF (20.4MB)
Morphometry and Relative Growth of Ophiolepis crassa (Echinodermata: Ophiuroidea), a Brittle Star from the Eastern Pacific
Laura Sanvicente-Añorve, Francisco A. Solís-Marín, and Itzel Rosales-Contreras (doi:10.6620/ZS.2021.60-26)
AbstractPDF (1.68MB)
Crustacean Fauna of the Aral Sea and its Relation to Ichthyofauna during the Modern Regression Crisis and Efforts at Restoration
Igor S. Plotnikov, Nikolai V. Aladin, Jens Mossin, and Jens T. Høeg (doi:10.6620/ZS.2021.60-25)
AbstractPDF (14.9MB)
Two New Moray Eels of Genera Diaphenchelys and Gymnothorax (Anguilliformes: Muraenidae) from Taiwan and the Philippines
Wen-Chien Huang, David G. Smith, Kar-Hoe Loh, and Te-Yu Liao (doi:10.6620/ZS.2021.60-24)
AbstractPDF (10.9MB)
An Endemic and Endangered New Species of the Lizard Liolaemus montanus Group from Southwestern Peru (Iguania: Liolaemidae), with a Key for the Species of the L. reichei Clade
Aarón J. Quiroz, Ling Huamaní-Valderrama, Roberto C. Gutiérrez, Álvaro J. Aguilar-Kirigin, Evaristo López-Tejeda, Ana Lazo-Rivera, Wilson Huanca-Mamani, Pablo Valladares-Faúndez, Juan J. Morrone, José Cerdeña, Juan C. Chaparro, and Cristian S. Abdala (doi:10.6620/ZS.2021.60-23)
AbstractPDF (22.8MB)
Free-living Microturbellarians (Platyhelminthes) from Wetlands in Southern Brazil, with the Description of Three New Species
Jhoe Reyes, Daniela Binow, Rogério T. Vianna, Francisco Brusa, and Samantha E. Martins (doi:10.6620/ZS.2021.60-22)
AbstractPDF (32MB)
Two New Species of the Brazilian Millipede Genus Rhicnosthetus Hoffman, 2006 (Polydesmida: Chelodesmidae), Including a Key to Males
Rodrigo Salvador Bouzan, Luiz Felipe Moretti Iniesta, Antonio Domingos Brescovit (doi:10.6620/ZS.2021.60-21)
AbstractPDF (1.89MB)
Vocalization development in Geoffroy’s bat, Myotis emarginatus (Chiroptera: Vespertilionidae)
Robab Mehdizadeh, Hojjat Eghbali, and Mozafar Sharifi (doi:10.6620/ZS.2021.60-20)
AbstractPDF (1.89MB)
Morphological and Genetic Variation Among Populations of the Fiddler Crab Minuca burgersi (Holthuis, 1967) (Crustacea: Brachyura: Ocypodidae) from Shores of the Caribbean Basin and Western South Atlantic Ocean
C. L. Thurman, R. E. Alber, M. J. Hopkins, and Hsi-Te Shih (doi:10.6620/ZS.2021.60-19)
AbstractPDF (7.38MB)
Seasonal Changes in the Sexual Size Dimorphisms of the Chelipeds and Pleons of the Porcellanid Crab Petrolisthes japonicus
Katsuyuki Hamasaki and Shigeki Dan (doi:10.6620/ZS.2021.60-18)
AbstractPDF (592KB)
Checklist of Pseudoscorpions (Arachnida, Pseudoscorpiones) of Albania
Martina Červená, Giulio Gardini, Daniel Jablonski, and Jana Christophoryová (doi:10.6620/ZS.2021.60-17)
AbstractPDF (8.20MB)
The Head of Fannia pusio (Fanniidae: Diptera) as A Novel Source of Morphometric Data for the Assessment of Variation along Geographic and Biological Lines
Yolanda Bravo-Pena, José Herrera-Russert, Elena Romera, and José Galián(doi:10.6620/ZS.2021.60-16)
AbstractPDF (2.83MB)
Revision of Majella Ortmann, 1893 (Crustacea: Brachyura: Majidae), with Description of Two New Species from the Indian Ocean
Peter K. L. Ng, Bee Yan, Lee, and Bertrand Richer de Forges (doi:10.6620/ZS.2021.60-15)
AbstractPDF (50.6MB)
Description of A New Genus and Two New Species of Freshwater Crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India
Smrithy Raj, Appukuttannair Biju Kumar, and Peter K. L. Ng (doi:10.6620/ZS.2021.60-14)
AbstractPDF (27.3MB)
A Taxonomic Review of the Buccal-attaching Fish Parasite Genus Lobothorax Bleeker, 1857 (Crustacea: Isopoda: Cymothoidae) with Description of A New Species from Southwestern India
Panakkool Thamban Aneesh, Niel L. Bruce, Appukuttannair Biju Kumar, M. Raj Bincy, and T. Mohanan Sreenath (doi10.6620/ZS.2021.60-13)
AbstractPDF (4.14MB)
Beetle Diversity Across Micro-habitats on Lizard Island Group (Great Barrier Reef, Australia)
John J. McCormack and Darko D. Cotoras (doi:10.6620/ZS.2021.60-12)
AbstractPDF (1.71MB)
Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species
Filipe Michels Bianchi, Verônica Krein, David Rider, and Jocelia Grazia (doi:10.6620/ZS.2021.60-11)
AbstractPDF (5.64MB)
A Taxonomic Review of the Gall Wasp Genus Saphonecrus Dalla-Torre and Kieffer and other Oak Cynipid Inquilines (Hymenoptera: Cynipidae) from Mainland China, with Updated Keys to Eastern Palaearctic and Oriental Species
Irene Lobato-Vila, Yiping Wang, George Melika, Rui Guo, Xiaoxue Ju, and Juli Pujade-Villar (doi:10.6620/ZS.2021.60-10)
AbstractPDF (3.70MB)
A Different Perspective on Sex Dimorphism in the Adult Hermann’s Tortoise: Geometric Morphometry
Sokol Duro, Ozan Gündemir, Bektaş Sönmez, Tefik Jashari, Tomasz Szara, Gülsün Pazvant, Arben Kambo  (doi:10.6620/ZS.2021.60-09)
AbstractPDF (2.12MB)
Mitochondrial Markers Identify a Genetic Boundary of the Green Tiger Prawn (Penaeus semisulcatus) in the Indo-Pacific Ocean
Siti Amalia Aisyah Abdul Halim, Ahmad Sofiman Othman, Noor Adelyna Mohammed Akib, Noorul-Azliana Jamaludin, Yuzine Esa, and Siti Azizah Mohd Nor  (doi:10.6620/ZS.2021.60-08)
AbstractPDF (2.79MB)
Revisiting a 135-year-old Taxonomic Account of the Freshwater Snail Semisulcospira multigranosa: Designating its Lectotype and Describing a New Species of the Genus (Mollusca: Gastropoda: Semisulcospiridae)
Naoto Sawada and Takafumi Nakano (doi:10.6620/ZS.2021.60-07)
AbstractPDF (16.8MB)
Differential Protein Metabolism and Regeneration in Gastrocnemius Muscles in High-fat Diet Fed Mice and Pre-hibernation Daurian Ground Squirrels: A Comparison between Pathological and Healthy Obesity
Xia Yan, Qiaohua Niu, Xuli Gao, Shenyang Shen, Nan He, Huiping Wang, Rongrong Fang, Yunfang Gao, and Hui Chang (doi:10.6620/ZS.2021.60-06)
AbstractPDF (9.78MB)
Nesting Biology of the Solitary Wasp Pisoxylon amenkei (Hymenoptera, Crabronidae, Trypoxylini) in a Neotropical Hotspot of Southern Brazil
Jean Pablo Alves de Deus, Caroline Nepomuceno Queiros, and Maria Luisa Tunes Buschini (doi:10.6620/ZS.2021.60-05)
AbstractPDF (6.85MB)
A Review of the Parasitic Isopod Genus Epicepon Nierstrasz & Brender à Brandis, 1931 (Crustacea: Isopoda: Epicaridea), Parasites of Cyclodorippoid Crabs (Crustacea: Decapoda: Brachyura), with Description of a New Species from New Caledonia
Christopher B. Boyko and Jason D. Williams (doi:10.6620/ZS.2021.60-04)
AbstractPDF (3.01MB)
Synchrony between Reproductive Phase and Flood Period: A Dispersion Mechanism for the Freshwater Clam Corbicula fluminea (Bivalvia: Corbiculidae) in a Brazilian Neotropical Floodplain
Claudenice Dei Tos,  Camila Antoniassi Cardim, Vanessa de Brito Pereira, and Roger Paulo Mormul (doi:10.6620/ZS.2021.60-03)
AbstractPDF (7.59MB)
Preliminary Study of Temperature Effects on Size and Shape in the Modern Spinicaudatan Eulimnadia texana (Crustacea: Branchiopoda)
Manja Hethke, Stephen C. Weeks, Veronika Schöttle, and D. Christopher Rogers (doi:10.6620/ZS.2021.60-02)
AbstractPDF (10.2MB)
Two New Species and a Further Country Record of the Caridean Shrimp Genus Periclimenaeus Borradaile, 1915 from Korea (Decapoda: Palaemonidae)
Jin-Ho Park and Sammy De Grave (doi:10.6620/ZS.2021.60-01)
AbstractPDF (19.4MB)