We publish high quality peer-reviewed articles on animal behavior, comparative physiology, evolution, ecology, and systematics and biogeography on animals taxa from all parts of the world.
OPEN ACCESS (Free Page Charge)
Zoological Studies is an OPEN ACCESS international journal and FREE OF PAGE CHARGES from authors.

Edited by:
Benny K.K. Chan (Editor-in-Chief)
Sen-Lin Tang (Co-Editor-in-Chief)

Impact Factor: 1.257
ISI Journal Citation Reports
Ranking: 2019: 78/168 (Zoology)

ISSN (E): 1810-522X
Follow us on Twitter @ZooStudies and Facebook

Zoological Studies is listed in PubMed
Sumbit manuscript

Zoological Studies Editorial Team

The Editor in Chief, Dr. Benny K.K. Chan, and the Co-editors in Chief, Dr. Sen-Lin Tang are professors/ research fellows at Biodiversity Research Center, Academia Sinica, Taiwan.

The editors and co-editors in chief are supported by 30 Associate Editors including Drs.Sheng-Feng Shen, I-Min Tso, Pung-Pung Hwang, Yeong-Choy Kam, David Miller, Shou-Hsien Li, Chung-Pin Lin, John Wang, Claudia Castellani, Yasuyuki Hashiguchi, Chih-Ming Hung, Chiun-Cheng Ko, Chi-Chien Kuo, Teng-Chiu Lin, Jian-Nan Liu, Ryuji Machida, Hiroomi Miyamoto, Yoko Nozawa, Gray A. Williams, Benny K.K. Chan, Allen Chen, Ka Hou Chu, Jens Thorvald Høeg, Jen-Pan Huang, Hin-Kiu Mok, Felipe Polivanov Ottoni, James D. Reimer, Shen-Horn Yen, Sen-Lin Tang, and Daryi Wang.

(See Editorial Board for details)

Message from the Editor in Chief

We take up a responsibility for the academic society

    Nowadays, we often need to pay expensive page charges/open-access fees to publish our works as most of the journals are now run by business publishers. Is it possible there are quality assured, peer-reviewed and open-access SCI scientific journals, in which colleagues can publish their research achievement free? We, Biodiversity Research Center, Academia Sinica, Taiwan, attempts to take up this responsibility. We run Zoological Studies, to provide a free SCI open-access publication platform for colleagues to share out their research achievements.
    Publish articles free of page charges does not mean Zoological Studies is lack of good quality. All the accepted articles passed a formal review system. Impact factors is an indication for the article citations in a journals and has implications for journal quality. However, article citations can be also affected by the ways of marketing methods for journals in commercial publishers. We are trying hard to maintain a good impact factor for Zoological Studies. All articles are 1) formally applied for a DOI number and being open-access in the web, 2) listed and housed in PubMed database and 3) promoted in our twitters and facebook. We are not under any umbrella of commercial publishers. All the editors and reviewers are volunteers for the academics. We hope you can support us to submit your quality works to Zoological Studies, and upload your published articles in Research Gate to enhance citations as well.

Prof. Benny K.K. Chan

Recent published articles in 2021
Ecological Role of Cyprideis torosa and Heterocypris salina (Crustacea, Ostracoda) in Saline Rivers of the Lake Elton Basin: Abundance, Biomass, Production, Fatty Acids
Vladimir A Gusakov, Olesia N Makhutova, Michail I Gladyshev, Larisa V Golovatyuk, and Tatiana D Zinchenko (doi:
Abstract丨PDF(-MB)(Proof production in process)
Threats to Site Occupation of Carnivores: A Spatiotemporal Encroachment of Non-native Species on the Native Carnivore Community in A Human-dominated Protected Area
Kasereka Vitekere, Luc Mumbere Lango, Jiao Wang, Mengyan Zhu, Guangshun Jiang, and Yan Hua (doi:
Abstract丨PDF(-MB)(Proof production in process)
Influence of Visitors on the Time Budget, Ranging and Strata Use of Lowe’s Monkey (Cercopithecus lowei) at Boabeng-Fiema Monkey Sanctuary, Ghana
Núria Badiella-Giménez, Bright Obeng Kankam, and Llorenç Badiella (doi:
Abstract丨PDF(-MB)(Proof production in process)
Taxonomy, Evolutionary and Dispersal Events of Pig-Tailed Macaque, Macaca nemestrina (Linnaeus, 1766) in Southeast Asia With Description of a New Subspecies, Macaca nemestrina perakensis in Malaysia
Muhammad Abu Bakar Abdul-Latiff and Badrul Munir Md-Zain (doi:
Abstract丨PDF(-MB)(Proof production in process)
Reproduction and Early Juvenile Growth of the Giant Clams Tridacna noae and Tridacna maxima in Taiwan
Po-Wei Su, Gwo-Liang Zhang, Po-Nien Chen, Keryea Soong, and Li-Lian Liu (doi:
Abstract丨PDF(-MB)(Proof production in process)
Ecomorphological Adaptations of Second Maxilliped-setation: Insights from Three Species of Fiddler Crabs from Panama
Shirley S.L. Lim and Sarah J.A. Goh (doi:
Abstract丨PDF(-MB)(Proof production in process)
Classification and Morphology of Rhinocypha spp. (Odonata): A Comprehensive Taxonomic Study within the Females
Mamat-Noorhidayah, Arpah-Abu, and Y. Norma-Rashid (doi:
Abstract丨PDF(-MB)(Proof production in process)
Moderate Population Structure in Drosophila sturtevanti from the South American Atlantic Forest Biome
Bruna Memari Trava, Rogério Pincela Mateus, Luciana Paes de Barros Machado, and Lilian Madi-Ravazzi (doi:
Abstract丨PDF(-MB)(Proof production in process)
Larval Performance of Amphidromous and Landlocked Atyid Shrimp Species in the Genus Paratya under Different Temperature and Salinity Conditions
Katsuyuki Hamasaki, Shuji Kondo, and Shigeki Dan (doi:
AbstractEarly view (551KB)
Thermal Physiological Performance and Thermal Metabolic Scope of the Whelk Kelletia kelletii (Forbes, 1850) (Gastropoda: Neptuneidae) Acclimated to Different Temperatures
Fernando Díaz, Ana Denise Re-Araujo, Eugenio Carpizo-Ituarte, Zaul Garcia-Esquivel, Ernesto Larios-Soriano, Leonel Perez-Carrasco, and Ernesto Lerma (doi:
AbstractEarly view (613KB)
Contact Reactions Between Individuals of the Coral-killing Sponge, Terpios hoshinota
Yurika Hirose, Siti Nurul Aini, and Hideyuki Yamashiro (doi:
AbstractEarly view (1.13MB)
Phylogenetic Position of Trichomycterus payaya and Examination of Osteological Characters Diagnosing the Neotropical Catfish Genus Ituglanis (Siluriformes: Trichomycteridae)
Wilson J. E. M. Costaa, José Leonardo O. Mattos, and Axel M. Katza (doi:
AbstractEarly view (2.90MB)
From Asia to Europe? Where Could the Geographical Place of Origin of Polymerus (Pachycentrum) carpathicus (Horváth, 1882) (Hemiptera: Miridae) Be?
Agnieszka Bugaj-Nawrocka, Artur Taszakowski, and Grzegorz Gierlasiński (doi:
AbstractEarly view (1.03MB)
Larval Morphology of Pit-building Antlions of the Tribe Myrmeleontini (Neuroptera, Myrmeleontidae) from Taiwan
Yu-Hsiu Lin, Jhih-Rong Liao, and Chiun-Cheng Ko (doi:
AbstractEarly view (4.45MB)
Seasonal Growth of the Purple Sea Urchin Heliocidaris crassispina Revealed by Sequential Fluorochrome Tagging
Juan Diego Urriago Suarez, Jane C.Y. Wong, Gilbert Lui, Clement P. Dumont, Jian-Wen Qiu, and Monthon Ganmanee (doi:
AbstractEarly view (841KB)
Identifying A New Phylogeographic Population of the Blyth’s Tragopan (Tragopan blythii) through Multi-locus Analyses
Jingyi Zou, Lu Dong, Geoffrey Davison, Win Hlaing, Moe Myint Aung, Yanyun Zhang, Zhengwang Zhang, Nan Wang, and De Chen (doi:
AbstractEarly view (793KB)
Plasticity of Foraging Strategies Adopted by the Painted Ghost Crab, Ocypode gaudichaudii, in Response to in situ Food Resource Manipulation Experiments
Adeline Y.P. Yong and Shirley S.L. Lim (doi:
AbstractEarly view (1.05MB)
Otolith Morphology: A Hidden Tool in the Taxonomic Study of Goatfishes (Teleostei: Perciformes: Mullidae)
Sorour Echreshavi, Hamid Reza Esmaeili, Azad Teimori, and Mohsen Safaie (doi:
AbstractEarly view (1.95MB)
Review of Chilean Cyrtinae (Diptera: Acroceridae) with the Description of Three New Species and the First Record of Villalus inanis from Argentina
Rodrigo M. Barahona-Segovia, Vicente Valdés Guzmán, and Laura Pañinao-Monsálvez (doi:-)
AbstractEarly view (1.68MB)
Littoral water in Hong Kong as a Potential Transient Habitat for Juveniles of a Temperate Deepwater Gnomefish, Scombrops boops (Acropomatiformes: Scombropidae)
Jiehong Wei, Jiarui Gu, Min Liu, Bai-An Lin, Gabriel Y. Lee, Tak-Cheung Wai, Paul K.S. Lam, Meng Yan, and Priscilla T.Y. Leung (doi:-)
AbstractEarly view (1.19MB)
Description of a New Species of the Marine Flatworm Prosthiostomum (Platyhelminthes: Polycladida) and its Three Known Congeners from Misaki, Japan, with Inference of Their Phylogenetic Positions within Prosthiostomidae
Aoi Tsuyuki, Hisanori Kohtsuka, and Hiroshi Kajihara (doi:-)
AbstractEarly view (2.01MB)
Spatial Distribution and Dietary Niche Breadth of Leopard Cat (Prionailurus bengalensis) Inhabiting Margalla Hills National Park, Pakistan
Victor Moctezuma and Gonzalo Halffter (doi:-)
AbstractEarly view (1.20MB)
Population Genetics Analysis of a Pomacea Snail (Gastropoda: Ampullariidae) in Thailand and its Low Infection by Angiostrongylus cantonensis
Abdulhakam Dumidae, Pichamon Janthu, Chanakan Subkrasae, Raxsina Polseela, Bandid Mangkit, Aunchalee Thanwisai, and Apichat Vitta (doi:-)
AbstractEarly view (2.11MB)
Species Redescriptions and New Species of the Onthophagus mexicanus Species Group (Coleoptera: Scarabaeidae), with Notes on Distribution and Rodent-dung Beetle Associations
Victor Moctezuma and Gonzalo Halffter (doi:-)
AbstractEarly view (8.29MB)
Setteleia, a New Genus for Four New Species from the Philippines (Insecta: Lepidoptera: Erebidae: Arctiinae)
Anton V. Volynkin and Karel Černý (doi:-)
AbstractEarly view (3.57MB)
Tissue-specific Isotopic Incorporation Turnover Rates and Trophic Discrimination Factors in the Freshwater Shrimp Macrobrachium borellii (Crustacea: Decapoda: Palaemonidae)
M. Florencia Viozzi, Carlos Martínez del Rio, and Verónica Williner (doi:-)
AbstractEarly view (503KB)
Taxonomic Revision of Living Pholadidae Lamarck, 1809 (Bivalvia: Myida) from South Western Atlantic
Marina Delfino and Javier H. Signorelli (doi:-)
AbstractEarly view (2.29MB)
AbstractPDF (1.68MB)
Crustacean Fauna of the Aral Sea and its Relation to Ichthyofauna during the Modern Regression Crisis and Efforts at Restoration
Igor S. Plotnikov, Nikolai V. Aladin, Jens Mossin, and Jens T. Høeg (doi:10.6620/ZS.2021.60-25)
AbstractPDF (14.9MB)
Two New Moray Eels of Genera Diaphenchelys and Gymnothorax (Anguilliformes: Muraenidae) from Taiwan and the Philippines
Wen-Chien Huang, David G. Smith, Kar-Hoe Loh, and Te-Yu Liao (doi:10.6620/ZS.2021.60-24)
AbstractPDF (10.9MB)
An Endemic and Endangered New Species of the Lizard Liolaemus montanus Group from Southwestern Peru (Iguania: Liolaemidae), with a Key for the Species of the L. reichei Clade
Aarón J. Quiroz, Ling Huamaní-Valderrama, Roberto C. Gutiérrez, Álvaro J. Aguilar-Kirigin, Evaristo López-Tejeda, Ana Lazo-Rivera, Wilson Huanca-Mamani, Pablo Valladares-Faúndez, Juan J. Morrone, José Cerdeña, Juan C. Chaparro, and Cristian S. Abdala (doi:10.6620/ZS.2021.60-23)
AbstractPDF (22.8MB)
Free-living Microturbellarians (Platyhelminthes) from Wetlands in Southern Brazil, with the Description of Three New Species
Jhoe Reyes, Daniela Binow, Rogério T. Vianna, Francisco Brusa, and Samantha E. Martins (doi:10.6620/ZS.2021.60-22)
AbstractPDF (32MB)
Two New Species of the Brazilian Millipede Genus Rhicnosthetus Hoffman, 2006 (Polydesmida: Chelodesmidae), Including a Key to Males
Rodrigo Salvador Bouzan, Luiz Felipe Moretti Iniesta, Antonio Domingos Brescovit (doi:10.6620/ZS.2021.60-21)
AbstractPDF (1.89MB)
Vocalization development in Geoffroy’s bat, Myotis emarginatus (Chiroptera: Vespertilionidae)
Robab Mehdizadeh, Hojjat Eghbali, and Mozafar Sharifi (doi:10.6620/ZS.2021.60-20)
AbstractPDF (1.89MB)
Morphological and Genetic Variation Among Populations of the Fiddler Crab Minuca burgersi (Holthuis, 1967) (Crustacea: Brachyura: Ocypodidae) from Shores of the Caribbean Basin and Western South Atlantic Ocean
C. L. Thurman, R. E. Alber, M. J. Hopkins, and Hsi-Te Shih (doi:10.6620/ZS.2021.60-19)
AbstractPDF (7.38MB)
Seasonal Changes in the Sexual Size Dimorphisms of the Chelipeds and Pleons of the Porcellanid Crab Petrolisthes japonicus
Katsuyuki Hamasaki and Shigeki Dan (doi:10.6620/ZS.2021.60-18)
AbstractPDF (592KB)
Checklist of Pseudoscorpions (Arachnida, Pseudoscorpiones) of Albania
Martina Červená, Giulio Gardini, Daniel Jablonski, and Jana Christophoryová (doi:10.6620/ZS.2021.60-17)
AbstractPDF (8.20MB)
The Head of Fannia pusio (Fanniidae: Diptera) as A Novel Source of Morphometric Data for the Assessment of Variation along Geographic and Biological Lines
Yolanda Bravo-Pena, José Herrera-Russert, Elena Romera, and José Galián(doi:10.6620/ZS.2021.60-16)
AbstractPDF (2.83MB)
Revision of Majella Ortmann, 1893 (Crustacea: Brachyura: Majidae), with Description of Two New Species from the Indian Ocean
Peter K. L. Ng, Bee Yan, Lee, and Bertrand Richer de Forges (doi:10.6620/ZS.2021.60-15)
AbstractPDF (50.6MB)
Description of A New Genus and Two New Species of Freshwater Crab (Decapoda: Brachyura: Gecarcinucidae) from the Southwestern Ghats, India
Smrithy Raj, Appukuttannair Biju Kumar, and Peter K. L. Ng (doi:10.6620/ZS.2021.60-14)
AbstractPDF (27.3MB)
A Taxonomic Review of the Buccal-attaching Fish Parasite Genus Lobothorax Bleeker, 1857 (Crustacea: Isopoda: Cymothoidae) with Description of A New Species from Southwestern India
Panakkool Thamban Aneesh, Niel L. Bruce, Appukuttannair Biju Kumar, M. Raj Bincy, and T. Mohanan Sreenath (doi10.6620/ZS.2021.60-13)
AbstractPDF (4.14MB)
Beetle Diversity Across Micro-habitats on Lizard Island Group (Great Barrier Reef, Australia)
John J. McCormack and Darko D. Cotoras (doi:10.6620/ZS.2021.60-12)
AbstractPDF (1.71MB)
Finding Padaeus bovillus (Hemiptera: Pentatomidae): A Phylogenetic Placement and the Description of Its Sister Species
Filipe Michels Bianchi, Verônica Krein, David Rider, and Jocelia Grazia (doi:10.6620/ZS.2021.60-11)
AbstractPDF (5.64MB)
A Taxonomic Review of the Gall Wasp Genus Saphonecrus Dalla-Torre and Kieffer and other Oak Cynipid Inquilines (Hymenoptera: Cynipidae) from Mainland China, with Updated Keys to Eastern Palaearctic and Oriental Species
Irene Lobato-Vila, Yiping Wang, George Melika, Rui Guo, Xiaoxue Ju, and Juli Pujade-Villar (doi:10.6620/ZS.2021.60-10)
AbstractPDF (3.70MB)
A Different Perspective on Sex Dimorphism in the Adult Hermann’s Tortoise: Geometric Morphometry
Sokol Duro, Ozan Gündemir, Bektaş Sönmez, Tefik Jashari, Tomasz Szara, Gülsün Pazvant, Arben Kambo  (doi:10.6620/ZS.2021.60-09)
AbstractPDF (2.12MB)
Mitochondrial Markers Identify a Genetic Boundary of the Green Tiger Prawn (Penaeus semisulcatus) in the Indo-Pacific Ocean
Siti Amalia Aisyah Abdul Halim, Ahmad Sofiman Othman, Noor Adelyna Mohammed Akib, Noorul-Azliana Jamaludin, Yuzine Esa, and Siti Azizah Mohd Nor  (doi:10.6620/ZS.2021.60-08)
AbstractPDF (2.79MB)
Revisiting a 135-year-old Taxonomic Account of the Freshwater Snail Semisulcospira multigranosa: Designating its Lectotype and Describing a New Species of the Genus (Mollusca: Gastropoda: Semisulcospiridae)
Naoto Sawada and Takafumi Nakano (doi:10.6620/ZS.2021.60-07)
AbstractPDF (16.8MB)
Differential Protein Metabolism and Regeneration in Gastrocnemius Muscles in High-fat Diet Fed Mice and Pre-hibernation Daurian Ground Squirrels: A Comparison between Pathological and Healthy Obesity
Xia Yan, Qiaohua Niu, Xuli Gao, Shenyang Shen, Nan He, Huiping Wang, Rongrong Fang, Yunfang Gao, and Hui Chang (doi:10.6620/ZS.2021.60-06)
AbstractPDF (9.78MB)
Nesting Biology of the Solitary Wasp Pisoxylon amenkei (Hymenoptera, Crabronidae, Trypoxylini) in a Neotropical Hotspot of Southern Brazil
Jean Pablo Alves de Deus, Caroline Nepomuceno Queiros, and Maria Luisa Tunes Buschini (doi:10.6620/ZS.2021.60-05)
AbstractPDF (6.85MB)
A Review of the Parasitic Isopod Genus Epicepon Nierstrasz & Brender à Brandis, 1931 (Crustacea: Isopoda: Epicaridea), Parasites of Cyclodorippoid Crabs (Crustacea: Decapoda: Brachyura), with Description of a New Species from New Caledonia
Christopher B. Boyko and Jason D. Williams (doi:10.6620/ZS.2021.60-04)
AbstractPDF (3.01MB)
Synchrony between Reproductive Phase and Flood Period: A Dispersion Mechanism for the Freshwater Clam Corbicula fluminea (Bivalvia: Corbiculidae) in a Brazilian Neotropical Floodplain
Claudenice Dei Tos,  Camila Antoniassi Cardim, Vanessa de Brito Pereira, and Roger Paulo Mormul (doi:10.6620/ZS.2021.60-03)
AbstractPDF (7.59MB)
Preliminary Study of Temperature Effects on Size and Shape in the Modern Spinicaudatan Eulimnadia texana (Crustacea: Branchiopoda)
Manja Hethke, Stephen C. Weeks, Veronika Schöttle, and D. Christopher Rogers (doi:10.6620/ZS.2021.60-02)
AbstractPDF (10.2MB)
Two New Species and a Further Country Record of the Caridean Shrimp Genus Periclimenaeus Borradaile, 1915 from Korea (Decapoda: Palaemonidae)
Jin-Ho Park and Sammy De Grave (doi:10.6620/ZS.2021.60-01)
AbstractPDF (19.4MB)