We publish high quality peer-reviewed articles on animal behavior, comparative physiology, evolution, ecology, and systematics and biogeography on animals taxa from all parts of the world.
OPEN ACCESS (Free Page Charge)
Zoological Studies is an OPEN ACCESS international journal and FREE OF PAGE CHARGES from authors.

Edited by:
Benny K.K. Chan (Editor-in-Chief)
Sen-Lin Tang (Co-Editor-in-Chief)

Impact Factor: 2.058
ISI Journal Citation Reports
Ranking: 2020: 53/175 (Zoology)

ISSN (E): 1810-522X
Follow us on Twitter @ZooStudies and Facebook

Zoological Studies is listed in PubMed
Sumbit manuscript

Zoological Studies Editorial Team

The Editor in Chief, Dr. Benny K.K. Chan, and the Co-editors in Chief, Dr. Sen-Lin Tang are professors/ research fellows at Biodiversity Research Center, Academia Sinica, Taiwan.

The editors and co-editors in chief are supported by 30 Associate Editors including Drs.Sheng-Feng Shen, I-Min Tso, Pung-Pung Hwang, Yeong-Choy Kam, David Miller, Shou-Hsien Li, Chung-Pin Lin, John Wang, Claudia Castellani, Yasuyuki Hashiguchi, Chih-Ming Hung, Chiun-Cheng Ko, Chi-Chien Kuo, Teng-Chiu Lin, Jian-Nan Liu, Ryuji Machida, Hiroomi Miyamoto, Yoko Nozawa, Gray A. Williams, Benny K.K. Chan, Allen Chen, Ka Hou Chu, Jens Thorvald Høeg, Jen-Pan Huang, Hin-Kiu Mok, Felipe Polivanov Ottoni, James D. Reimer, Shen-Horn Yen, Sen-Lin Tang, and Daryi Wang.

(See Editorial Board for details)

Message from the Editor in Chief

We take up a responsibility for the academic society

    Nowadays, we often need to pay expensive page charges/open-access fees to publish our works as most of the journals are now run by business publishers. Is it possible there are quality assured, peer-reviewed and open-access SCI scientific journals, in which colleagues can publish their research achievement free? We, Biodiversity Research Center, Academia Sinica, Taiwan, attempts to take up this responsibility. We run Zoological Studies, to provide a free SCI open-access publication platform for colleagues to share out their research achievements.
    Publish articles free of page charges does not mean Zoological Studies is lack of good quality. All the accepted articles passed a formal review system. Impact factors is an indication for the article citations in a journals and has implications for journal quality. However, article citations can be also affected by the ways of marketing methods for journals in commercial publishers. We are trying hard to maintain a good impact factor for Zoological Studies. All articles are 1) formally applied for a DOI number and being open-access in the web, 2) listed and housed in PubMed database and 3) promoted in our twitters and facebook. We are not under any umbrella of commercial publishers. All the editors and reviewers are volunteers for the academics. We hope you can support us to submit your quality works to Zoological Studies, and upload your published articles in Research Gate to enhance citations as well.

Prof. Benny K.K. Chan

Recent published articles in 2021
Populational evidences support monogamy mating system in five species of snapping shrimps of the genus Alpheus (Caridea: Alpheidae)
Ana Carla Costa de Souza, José Roberto Souza, and Alexandre Oliveira Almeida (doi:
Abstract丨PDF(-MB)(Proof production in process)
Macroinvertebrate Communities and Body Condition of Larval Eastern Hellbender Salamanders (Cryptobranchus alleganiensis) in North Carolina
Shem Unger, Sierra Benfield, Lori Williams, Charles Lawson, and John Groves (doi:
Abstract丨PDF(-MB)(Proof production in process)
Metabarcoding of Fish Larvae in Merbok River Reveals Species Diversity and Distribution Along its Mangrove Environment
Norli Fauzani Mohd Abu Hassan Alshari, Siti Zuliana Ahmad, Azali Azlan, Lee Youn-Ho, Ghows Azzam, and Siti Azizah Mohd Nor (doi:
AbstractEarly view (574KB)
Molecular Identification of Mid to Final Stage Slipper Lobster Phyllosoma Larvae of the Genus Chelarctus (Crustacea: Decapoda: Scyllaridae) Collected in the Pacific with Descriptions of Their Larval Morphology
Kenta Ueda, Takashi Yanagimoto, Seinen Chow, Mari Kuroki, and Takashi Yamakawa (doi:
AbstractEarly view (2.05MB)
Tolerance to Anhydrobiotic Conditions Among Two Coexisting Tardigrade Species Differing in Life Strategies
Milena Roszkowska, Bartłomiej Gołdyn, Daria Wojciechowska, Jakub Z. Kosicki, Edyta Fiałkowska, Hanna Kmita, and Łukasz Kaczmarek (doi:
AbstractEarly view (0.97MB)
Multiple Environmental Factors Increase the Niche Complexity and Species Diversity of Brachyuran Crabs in an Intertidal Algal Reef Ecosystem in Northwestern Taiwan
Kun-Chang Li, Hung-Chang Liu, and Hui-Chen Lin (doi:
AbstractPDF (2.98MB)
A Soil Nematode Community Response to Reclamation of Salinized Abandoned Farmland
Lei Yang, Fenghua Zhang, and Yanqin Luo (doi:
AbstractEarly view (906KB)
Quantifying the Effect of Land-cover Change on the Endangered Farmland Green Treefrog (Zhangixalus arvalis) in an Agricultural Landscape: Implications for Conservation
Sin Chen, Meng-Hsien Chuang, Hau-Jie Shiu, and Jian-Nan Liu (doi:
AbstractEarly view (1.13MB)
High Mountain Echiniscid (Heterotardigrada) Fauna of Taiwan
Piotr Gąsiorek, Katarzyna Vončina, Reinhardt Møbjerg Kristensen, and Łukasz Michalczyk (doi:
AbstractEarly view (14.9MB)
Bacterial Communities in Multiple Tissues Across the Body Surface of Three Coastal Shark Species
Chelsea Black, Liza Merly, and Neil Hammerschlag (
AbstractPDF (7.70MB)
A New Species of the Intertidal Limpet Eoacmaea (Patellogastropoda: Eoacmaeidae) from Yonaguni Island, Japan and Taiwan
Tomoyuki Nakano, Takenori Sasaki, Takeharu Kosuge, and Benny K.K. Chan (doi:10.6620/ZS.2021.60-68)
AbstractPDF (8.48MB)
Dietary Habits of Free-ranging Banded Langur (Presbytis femoralis) in a Secondary-human Modified Forest in Johor, Malaysia
Mohd Faudzir Najmuddin, Hidayah Haris, Noratiqah Norazlimi, Farhani Ruslin, Ikki Matsuda, Badrul Munir Md-Zain, and Muhammad Abu Bakar Abdul-Latiff (doi:
AbstractPDF (1.54MB)
The Hillstream Decapod Crustaceans of Shenzhen, China, with Description of a New Species of Freshwater Crab (Crustacea: Brachyura: Potamidae) of the Genus Megapleonum Huang, Shih & Ahyong, 2018
Chao Huang and Si-Ying Mao (doi:
AbstractPDF (14.9MB)
A New Species of the Anchovy Genus Stolephorus Lacepède 1803 from North Sumatra, Indonesia, and Redescriptions of Stolephorus pacificus Baldwin 1984 and Stolephorus teguhi Kimura, Hori and Shibukawa 2009 (Teleostei: Clupeiformes: Engraulidae)
Harutaka Hata and Hiroyuki Motomura (
AbstractPDF (17.9MB)
Acoustic Properties of the Otolith of the Large Yellow Croaker Larimichthys crocea (Perciformes: Sciaenidae)
Xin-Hai Zhang, Yi Tao, Yang-Liang Zhou, Li-Guo Tang, Min Liu, and Xiao-Mei Xu (doi:
AbstractPDF (6.57MB)
Population Genetic Structures of the Jellyfish Aurelia coerulea Polyps on Korean Coasts and Implications as Revealed by Mitochondrial COI
Yoseph Seo, Buhari Lawan Muhammad, Jinho Chae, and Jang-Seu Ki
AbstractPDF (2.29MB)
Life History Traits of the Snapping Shrimp Alpheus carlae (Decapoda: Alpheidae) from the South-eastern Coast of Brazil
Yoseph Seo, Buhari Lawan Muhammad, Jinho Chae, and Jang-Seu Ki (doi:
AbstractPDF (33.1MB)
Histological Evidence of Multiple Spawning in Wild Female Japanese Eel Anguilla japonica
Akio Shimizu, Shigeho Ijiri, Hikari Izumi, Koichiro Gen, Hiroaki Kurogi, Hiroshi Hashimoto, Hideki Tanaka, Tadao Jinbo, Hiroaki Saito, and Seinen Chow (doi:
AbstractPDF (6.75MB)
Diversity in the Taiwanese Swimming Crabs (Crustacea: Brachyura: Portunidae) Estimated through DNA Barcodes, with Descriptions of 14 New Records
Yu-Hsuan Huang and Hsi-Te Shih (doi:
AbstractPDF (25.0MB)
A New Species of the Genus Ophiomonas Djakonov (Echinodermata: Ophiuroidea: Amphilepididae) from the Deep-Sea of Japan
Masanori Okanishi, Takumi Matsuo, and Toshihiko Fujita (doi:10.6620/ZS.2021.60-59)
AbstractPDF (37.1MB)
Preliminary Observations on the Effect of Temperature and Food Concentration on the Egg Production Rate and Hatching Success of Acartia amboinensis from the Central Red Sea
Mohsen M. El-Sherbiny and Ali Al-Aidaroos
AbstractPDF (1.86MB)
A New Species of Amphipod Pariphinotus Kunkel, 1910 (Amphipoda: Phliantidae) from Southwestern Atlantic
André R. Senna, Urssula N. Guedes, Luiz F. Andrade, and Guilherme H. Pereira-Filho
AbstractPDF (7.79MB)
Effects of Farming Systems on Insect Communities in the Paddy Fields of a Simplified Landscape During a Pest-control Intervention
Chao-Nien Koh, Ming-Chih Chiu, Ling-Mu Jaung, Yung-Jen Lu, and
and Hsing-Juh Lin (doi:10.6620/ZS.2021.60-56)
AbstractPDF (2.14MB)
Diversity and Species Composition of Bark and Ambrosia Beetles Captured Using Ethanol Baited Traps on Different Hosts in East Java, Indonesia
Hagus Tarno, Yogo Setiawan, Cindy Budi Kusuma, Miftachul Fitriyah, Ahmad Hudan Nazaruddin, Alvian Putra Yawandika, Hanif Ardiansyah Nasution, Ronauli Saragih, Achmad Praditya Yoga Bagasta, Zheng Wang, and Jianguo
Wang (doi:10.6620/ZS.2021.60-55)
AbstractPDF (4.81MB)
Sympatric Two-species Infestation by Rhizocephalan Barnacle Parasites in the Spider Crab Pugettia aff. ferox Ohtsuchi & Kawamura, 2019 from Peter the Great Bay (Northwestern Sea of Japan)
Darya D. Golubinskaya, Olga M. Korn, Svetlana N. Sharina, and Nikolai I. Selin (doi:
AbstractPDF (8.36MB)
Ecological Role of Cyprideis torosa and Heterocypris salina (Crustacea, Ostracoda) in Saline Rivers of the Lake Elton Basin: Abundance, Biomass, Production, Fatty Acids
ladimir A.Gusakov, Olesia N. Makhutova, Michail I. Gladyshev, Larisa V. Golovatyuk, and Tatiana D. Zinchenko (doi:10.6620/ZS.2021.60-53)
AbstractPDF (2.43MB)
Threats to Site Occupation of Carnivores: A Spatiotemporal Encroachment of Non-native Species on the Native Carnivore Community in A Human-dominated Protected Area
Kasereka Vitekere, Luc Mumbere Lango, Jiao Wang, Mengyan Zhu, Guangshun Jiang, and Yan Hua
AbstractPDF (3.16MB)
Influence of Visitors on the Time Budget, Ranging and Strata Use of Lowe’s Monkey (Cercopithecus lowei) at Boabeng-Fiema Monkey Sanctuary, Ghana
Núria Badiella-Giménez, Bright Obeng Kankam, and Llorenç Badiella
AbstractPDF (7.00MB)
Taxonomy, Evolutionary and Dispersal Events of Pig-Tailed Macaque, Macaca nemestrina (Linnaeus, 1766) in Southeast Asia With Description of a New Subspecies, Macaca nemestrina perakensis in Malaysia
Muhammad Abu Bakar Abdul-Latiff and Badrul Munir Md-Zain
AbstractPDF (7.94MB)
Reproduction and Early Juvenile Growth of the Giant Clams Tridacna noae and Tridacna maxima in Taiwan
Po-Wei Su, Gwo-Liang Zhang,
Bonien Chen, Keryea Soong, and Li-Lian Liu (doi:10.6620/ZS.2021.60-49)
AbstractPDF (2.10MB)
Ecomorphological Adaptations of Second Maxilliped-setation: Insights from Three Species of Fiddler Crabs from Panama
hirley S.L. Lim and Sarah J.A. Goh (doi:10.6620/ZS.2021.60-48)
AbstractPDF (4.04MB)
Classification and Morphology of Rhinocypha spp. (Odonata): A Comprehensive Taxonomic Study within the Females
oorhidayah Mamat, Arpah-Abu, and Y. Norma-Rashid (doi:10.6620/ZS.2021.60-47)
AbstractPDF (MB)
Moderate Population Structure in Drosophila sturtevanti from the South American Atlantic Forest Biome
runa Memari Trava, Rogério Pincela Mateus, Luciana Paes de Barros Machado, and Lilian Madi-Ravazzi (doi:10.6620/ZS.2021.60-46)
AbstractPDF (3.17MB)
Larval Performance of Amphidromous and Landlocked Atyid Shrimp Species in the Genus Paratya under Different Temperature and Salinity Conditions
atsuyuki Hamasaki, Shuji Kondo, and Shigeki Dan (doi:10.6620/ZS.2021.60-45)
AbstractPDF (7.35MB)
Thermal Physiological Performance and Thermal Metabolic Scope of the Whelk Kelletia kelletii (Forbes, 1850) (Gastropoda: Neptuneidae) Acclimated to Different Temperatures
Fernando Díaz, Ana Denise Re-Araujo, Eugenio Carpizo-Ituarte, Zaul Garcia-Esquivel, Ernesto Larios-Soriano, Leonel Perez-Carrasco, and Ernesto Lerma (doi:10.6620/ZS.2021.60-44)
AbstractPDF (2.09MB)
Phylogenetic Position of Trichomycterus payaya and Examination of Osteological Characters Diagnosing the Neotropical Catfish Genus Ituglanis (Siluriformes: Trichomycteridae)
Wilson J. E. M. Costaa, José Leonardo O. Mattos, and Axel M. Katza (doi:10.6620/ZS.2021.60-43)
AbstractPDF (7.88MB)
From Asia to Europe? Where Could the Geographical Place of Origin of Polymerus (Pachycentrum) carpathicus (Horváth, 1882) (Hemiptera: Miridae) Be?
Agnieszka Bugaj-Nawrocka, Artur Taszakowski, and Grzegorz Gierlasiński (doi:-10.6620/ZS.2021.60-42)
AbstractPDF (37.4MB)
Contact Reactions Between Individuals of the Coral-killing Sponge, Terpios hoshinota
urika Hirose, Siti Nurul Aini, and Hideyuki Yamashiro (doi:10.6620/ZS.2021.60-41)
AbstractPDF (3.23MB)
Identifying A New Phylogeographic Population of the Blyth’s Tragopan (Tragopan blythii) through Multi-locus Analyses
Jingyi Zou, Lu Dong, Geoffrey Davison, Win Hlaing, Moe Myint Aung, Yanyun Zhang, Zhengwang Zhang, Nan Wang, and De Chen (doi:10.6620/ZS.2021.60-40)
AbstractPDF (3.82MB)
Larval Morphology of Pit-building Antlions of the Tribe Myrmeleontini (Neuroptera, Myrmeleontidae) from Taiwan
Yu-Hsiu Lin, Jhih-Rong Liao, and Chiun-Cheng Ko (doi:10.6620/ZS.2021.60-39)
AbstractPDF (25.8MB)
Seasonal Growth of the Purple Sea Urchin Heliocidaris crassispina Revealed by Sequential Fluorochrome Tagging
Juan Diego Urriago Suarez, Jane C.Y. Wong, Gilbert Lui, Clement P. Dumont, Jian-Wen Qiu, and Monthon Ganmanee (doi:
AbstractPDF (12.6MB)
Plasticity of Foraging Strategies Adopted by the Painted Ghost Crab, Ocypode gaudichaudii, in Response to in situ Food Resource Manipulation Experiments
Adeline Y.P. Yong and Shirley S.L. Lim
AbstractPDF (7.98MB)
Otolith Morphology: A Hidden Tool in the Taxonomic Study of Goatfishes (Teleostei: Perciformes: Mullidae)
Sorour Echreshavi, Hamid Reza Esmaeili, Azad Teimori, and Mohsen Safaie (doi:
AbstractPDF (14.8MB)